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Santana - Crato, Ceará, Brazil

Formation	Santana
Member	Crato
Age:	122 - 112 Ma Early/Lower Cretaceous
Interval	Late Aptian
Lithology	lime mudstone
Number of Collections	3
Number of Occurrences	22

Recorded Sample Locations

Location	Region	Stratigraphic Name
Crato	Ceará, Brazil	Santana - Crato

Associated Units

Stratigraphic Name	Age	Lithology	Occurrence Records
Crato	122 - 112 Ma Early/Lower Cretaceous	lime mudstone, mudstone	4
Araripe - Crato	113 - 101 Ma Early/Lower Cretaceous	limestone, not reported/"limestone", "carbonate", lime mudstone	659
Araripe - Crato - Nova Olinda	122 - 109 Ma Early/Lower Cretaceous	lime mudstone/mudstone, lime mudstone, "limestone"	98

Recorded Fossils

Accepted Name	Type	Hierarchy	Age
Cretadiplura ceara species	7	Animalia : Arthropoda : Arachnida : Araneae : Dipluridae : Cretadiplura : *Cretadiplura ceara*	122.46 - 112.03 Ma Early/Lower Cretaceous
Dinodiplura ambulacra species	7	Animalia : Arthropoda : Arachnida : Araneae : Dipluridae : Dinodiplura : *Dinodiplura ambulacra*	122.46 - 112.03 Ma Early/Lower Cretaceous
Protoischnurus axelrodorum species	7	Animalia : Arthropoda : Arachnida : Scorpiones : Protoischnuridae : Protoischnurus : *Protoischnurus axelrodorum*	122.46 - 112.03 Ma Early/Lower Cretaceous
Scolopendromorpha order	4	Animalia : Arthropoda : Chilopoda : Scolopendromorpha	122.46 - 112.03 Ma Early/Lower Cretaceous
Blattodea order	4	Animalia : Arthropoda : Insecta : Blattodea	122.46 - 112.03 Ma Early/Lower Cretaceous
Cratovitisma oldreadi species	7	Animalia : Arthropoda : Insecta : Blattodea : Cratovitismidae : Cratovitisma : *Cratovitisma oldreadi*	122.46 - 112.03 Ma Early/Lower Cretaceous
Piniblattella limai species	7	Animalia : Arthropoda : Insecta : Blattodea : Ectobiidae : Piniblattella : *Piniblattella limai*	122.46 - 112.03 Ma Early/Lower Cretaceous
Ponopterix axelrodi species	7	Animalia : Arthropoda : Insecta : Blattodea : Umenocoleidae : Ponopterix : *Ponopterix axelrodi*	122.46 - 112.03 Ma Early/Lower Cretaceous
Coleoptera order	4	Animalia : Arthropoda : Insecta : Coleoptera	122.46 - 112.03 Ma Early/Lower Cretaceous
Diptera order	4	Animalia : Arthropoda : Insecta : Diptera	122.46 - 112.03 Ma Early/Lower Cretaceous
Ephemeroptera order	4	Animalia : Arthropoda : Insecta : Ephemeroptera	122.46 - 112.03 Ma Early/Lower Cretaceous
Hemiptera order	4	Animalia : Arthropoda : Insecta : Hemiptera	122.46 - 112.03 Ma Early/Lower Cretaceous
Christometra paradoxa species	7	Animalia : Arthropoda : Insecta : Hemiptera : Hydrometridae : Christometra : *Christometra paradoxa*	122.46 - 112.03 Ma Early/Lower Cretaceous
Apocrita suborder	4.3	Animalia : Arthropoda : Insecta : Hymenoptera : Apocrita	122.46 - 112.03 Ma Early/Lower Cretaceous
Dryellina placida species	7	Animalia : Arthropoda : Insecta : Neuroptera : Mesochrysopidae : Dryellina : *Dryellina placida*	122.46 - 112.03 Ma Early/Lower Cretaceous
Nuddsia repatriata species	7	Animalia : Arthropoda : Insecta : Neuroptera : Osmylidae : Nuddsia : *Nuddsia repatriata*	122.46 - 112.03 Ma Early/Lower Cretaceous
Orthoptera order	4	Animalia : Arthropoda : Insecta : Orthoptera	122.46 - 112.03 Ma Early/Lower Cretaceous
Cratoelcana zessini species	7	Animalia : Arthropoda : Insecta : Orthoptera : Elcanidae : Cratoelcana : *Cratoelcana zessini*	122.46 - 112.03 Ma Early/Lower Cretaceous
Elcanidae family	5	Animalia : Arthropoda : Insecta : Orthoptera : Elcanidae	122.46 - 112.03 Ma Early/Lower Cretaceous
Cratotetraspinus fossorius species	7	Animalia : Arthropoda : Insecta : Orthoptera : Gryllotalpidae : Cratotetraspinus : *Cratotetraspinus fossorius*	122.46 - 112.03 Ma Early/Lower Cretaceous

References

Menon F., Penney D., et al (2003) A new fossil scolopendromorph centipede from the Crato Formation of Brazil, Bulletin of the British Myriapod and Isopod Group 19, 62-66

Selden P. A., Casado F. C., et al (2006) Mygalomorph spiders (Araneae: Dipluridae) from the Lower Cretaceous Crato Lagerstätte, Araripe basin, north-east Brazil, Palaeontology 49, 817-826

Menon F. (2007) Higher systematics of scorpions from the Crato Formation, Lower Cretaceous of Brazil, Palaeontology 50, 185-195

Martins-Neto R. G., Rodrigues V. Z. (2009) New Neuroptera (Insecta, Osmylidae and Mesochrysopidae) from the Santana Formation, Lower Cretaceous of northeast Brazil, Gaea 5 1, 15-20

Martins-Neto R. G., Rodrigues V. Z. (2010) New neuropteran insects (Osmylidae, Palaeoleontidae, Araripeneuridae and Psychopsidae) from the Santana Formation, Early Cretaceous NE Brazil, Gaea 6, 1-8

Barling N., Martill D. M., et al (2015) High fidelity preservation of fossil insects from the Crato Formation (Lower Cretaceous) of Brazil, Cretaceous Research 52, 605-622

Pêgas R. V., Leal M. E. C., et al (2017) A surprisingly derived hydrometrid (Hemiptera, Gerromorpha) from the Crato Formation, Early Cretaceous of Brazil, Historical Biology 30, 1094-1101

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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