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Osmylidae

Photos

Photo Gallery

Description

Osmylidae are a small family of winged insects of the net-winged insect order Neuroptera. The osmylids, also called giant lacewings, are found all over the world. A common species through most of Europe is Osmylus fulvicephalus.

From Wikipedia article at https://en.wikipedia.org/wiki/Osmylidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	9380	https://www.gbif.org/species/9380
PaleoBioDB ID (PBDB)	115697	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=115697

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Neuroptera : Osmylidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
superorder	Neuropterida
-	Eidoneuroptera	Engel et al. 2018
order	Neuroptera	Linnaeus 1758
-	Euneuroptera	Engel et al. 2018
superfamily	Osmyloidea	Leach 1815
family	Osmylidae	Leach 1815

Common Name

lance lacewing

Scientific Name

Osmylidae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Osmylidae	family	belongs to Planipennia	stated without evidence	Riek, 1955
Osmylidae	family	belongs to Neuroptera	stated without evidence	Menon and Makarkin, 2008
Osmylidae	family	belongs to Osmyloidea	stated with evidence	Engel and Grimaldi, 2008

Status (PBDB)

extant

Taxon Size (PBDB)

113

Extant Size (PBDB)

9 (8.0%)

First Recorded Appearance

201 - 191 Ma
Early Jurassic

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

F. Menon and V. N. Makarkin. 2008. New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil. Palaeontology 51(1):149-162

Common Name(s)

Watergaasvliegen, Lance Lacewing, Vattenrovsländor

Wikipedia

https://en.wikipedia.org/wiki/Osmylidae

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Eidoporisminae subfamily	listed (PBDB)		1	136 Ma Early/Lower Cretaceous	0 Ma Extant
Gumillinae subfamily	listed (PBDB)		15	170 Ma Middle Jurassic	0 Ma Extant
Kempyninae subfamily	listed (PBDB)		17	183 Ma Early Jurassic	0 Ma Extant
Mesosmylininae ✝ subfamily	listed (PBDB)		9	201 Ma Early Jurassic	152 Ma Late Jurassic
Osmylinae subfamily	listed (PBDB)		3	37.2 Ma Eocene	0 Ma Extant
Protosmylinae subfamily	listed (PBDB)		17	201 Ma Early Jurassic	0 Ma Extant
Spilosmylinae subfamily	listed (PBDB)		2	166 Ma Middle Jurassic	0 Ma Extant
Allotriosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Arbusella ✝ genus	accepted (GBIF) listed (PBDB)		4	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Archaeosmylidia ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Australysmus genus	accepted (GBIF)	No associated record in PBDB
Bellinympha ✝ genus	accepted (GBIF) listed (PBDB)		2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Burmaleon genus	accepted (GBIF)	No associated record in PBDB
Carinosmylus genus	accepted (GBIF)	No associated record in PBDB
Clydosmylus genus	accepted (GBIF)	No associated record in PBDB
Cratosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	122 Ma Early/Lower Cretaceous	122 Ma Early/Lower Cretaceous
Cretosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Daohugosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Dimidiosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	152 Ma Late Jurassic	152 Ma Late Jurassic
Eidoporismus genus	accepted (GBIF)	No associated record in PBDB
Enodinympha ✝ genus	accepted (GBIF) listed (PBDB)		2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Ensiosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Epiosmylus ✝ genus	accepted (GBIF) listed (PBDB)		3	170 Ma Middle Jurassic	166 Ma Middle Jurassic
Erlikosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	191 Ma Early Jurassic	191 Ma Early Jurassic
Euosmylus genus	accepted (GBIF)	No associated record in PBDB
Euporismites ✝ genus	accepted (GBIF) listed (PBDB)		3	56.0 Ma Paleocene	56.0 Ma Paleocene
Euporismus genus	accepted (GBIF)	No associated record in PBDB
Glenosmylus genus	accepted (GBIF)	No associated record in PBDB
Grandosmylus genus	accepted (GBIF)	No associated record in PBDB
Gryposmylus genus	accepted (GBIF)	No associated record in PBDB
Gumilla genus	accepted (GBIF) listed (PBDB)				0 Ma Extant
Heterosmylus genus	accepted (GBIF)	No associated record in PBDB
Hyposmylus genus	accepted (GBIF)	No associated record in PBDB
Imanosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	72.1 Ma Late/Upper Cretaceous	72.1 Ma Late/Upper Cretaceous
Isostenosmylus genus	accepted (GBIF)	No associated record in PBDB
Juraheterosmylus ✝ genus	accepted (GBIF) listed (PBDB)		3	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Jurakempynus ✝ genus	accepted (GBIF) listed (PBDB)		5	166 Ma Middle Jurassic	152 Ma Late Jurassic
Jurosmylus ✝ genus	accepted (GBIF) listed (PBDB)		4	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Karaosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Kempynosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Kempynus genus	accepted (GBIF)	No associated record in PBDB
Kolbasinella ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Kubekius ✝ genus	accepted (GBIF) listed (PBDB)		1	170 Ma Middle Jurassic	170 Ma Middle Jurassic
Laccosmylus ✝ genus	accepted (GBIF) listed (PBDB)		4	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Lahulus genus	accepted (GBIF)	No associated record in PBDB
Lithosmylus ✝ genus	accepted (GBIF) listed (PBDB)		2	37.2 Ma Eocene	37.2 Ma Eocene
Lysmus genus	accepted (GBIF)	No associated record in PBDB
Mesosmylidus ✝ genus	accepted (GBIF) listed (PBDB)		1	145 Ma Late Jurassic	145 Ma Late Jurassic
Mesosmylina ✝ genus	accepted (GBIF) listed (PBDB)		7	191 Ma Early Jurassic	152 Ma Late Jurassic
Mesosmylus genus	accepted (GBIF)	No associated record in PBDB
Nilionympha ✝ genus	accepted (GBIF) listed (PBDB)		2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Nuddsia ✝ genus	accepted (GBIF) listed (PBDB)		3	122 Ma Early/Lower Cretaceous	99.6 Ma Late/Upper Cretaceous
Oedosmylus genus	accepted (GBIF)	No associated record in PBDB
Osmylidia ✝ genus	accepted (GBIF) listed (PBDB)		4	37.2 Ma Eocene	37.2 Ma Eocene
Osmylochrysa ✝ genus	accepted (GBIF) listed (PBDB)		2	145 Ma Late Jurassic	145 Ma Late Jurassic
Osmylopsis ✝ genus	accepted (GBIF) listed (PBDB)		1	145 Ma Late Jurassic	145 Ma Late Jurassic
Osmylus genus	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	0 Ma Extant
Palaeothyridosmylus ✝ genus	accepted (GBIF) listed (PBDB)		2	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Parosmylus genus	accepted (GBIF)	No associated record in PBDB
Paryphosmylus genus	accepted (GBIF)	No associated record in PBDB
Petrushevskia ✝ genus	accepted (GBIF) listed (PBDB)		2	201 Ma Early Jurassic	199 Ma Early Jurassic
Phlebosmylus genus	accepted (GBIF)	No associated record in PBDB
Phymatosmylus genus	accepted (GBIF)	No associated record in PBDB
Porismus genus	accepted (GBIF)	No associated record in PBDB
Pronymphites ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Protosmylina ✝ genus	accepted (GBIF) listed (PBDB)		1	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Protosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Pseudosmylidia ✝ genus	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Rudiosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Saucrosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Sauktangida ✝ genus	accepted (GBIF) listed (PBDB)		1	183 Ma Early Jurassic	183 Ma Early Jurassic
Scapoptera ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Sogjuta ✝ genus	accepted (GBIF) listed (PBDB)		2	201 Ma Early Jurassic	152 Ma Late Jurassic
Spilosmylus genus	accepted (GBIF)	No associated record in PBDB
Stenochrysa ✝ genus	accepted (GBIF) listed (PBDB)		1	145 Ma Late Jurassic	145 Ma Late Jurassic
Stenolysmus genus	accepted (GBIF)	No associated record in PBDB
Stenosmylina ✝ genus	accepted (GBIF) listed (PBDB)		1	136 Ma Early/Lower Cretaceous	136 Ma Early/Lower Cretaceous
Stenosmylus genus	accepted (GBIF)	No associated record in PBDB
Tengriosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	191 Ma Early Jurassic	191 Ma Early Jurassic
Tenuosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	166 Ma Middle Jurassic	166 Ma Middle Jurassic
Tetanoptilon ✝ genus	accepted (GBIF) listed (PBDB)		1	183 Ma Early Jurassic	183 Ma Early Jurassic
Thaumatosmylus genus	accepted (GBIF)	No associated record in PBDB
Thyridosmylus genus	accepted (GBIF)	No associated record in PBDB
Yanosmylus ✝ genus	accepted (GBIF) listed (PBDB)		1	125 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous

Synonymy List

Year	Name and Author
1815	Osmylidae Leach
1955	Osmylidae Riek p. 678
1992	Osmylidae Carpenter
2008	Osmylidae Engel and Grimaldi
2008	Osmylidae Menon and Makarkin
2018	Osmylidae Engel et al.
2019	Osmylidae Winterton et al. p. 13

References

Menon F., Makarkin V. N. (2008) New fossil lacewings and antlions (Insecta, Neuroptera) from the Lower Cretaceous Crato Formation of Brazil, Palaeontology 51 1, 149-162

Riek E. F. (1955) Fossil insects from the Triassic beds at Mt. Crosby, Queensland, Australian Journal of Zoology 3 4, 654-691

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655

Engel M. S., Grimaldi D. A. (2008) Diverse Neuropterida in Cretaceous amber, with particular reference to the paleofauna of Myanmar (Insecta), Nova Supplementa Entomologica 20, 1-86

Engel M. S., Winterton S. L., et al (2018) Phylogeny and evolution of Neuropterida: where have wings of lace taken us?, Annual Review of Entomology 63, 531-551

Winterton S. L., Martins C. C., et al (2019) Lance lacewings of the world (Neuroptera: Archeosmylidae, Osmylidae, Saucrosmylidae): review of living and fossil genera, Zootaxa 4581, 1-99

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera

Penny, Norman D., 1977: Lista de Megaloptera, Neuroptera e Raphidioptera do México, América Central, ilhas Caraíbas e América do Sul. Acta Amazonica, vol. 7, no. 4, Suplemento. 5-61. - via Integrated Taxonomic Information System (ITIS)

Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas

Meinander, M. (1996) - via Taxa Watermanagement the Netherlands (TWN)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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