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Empididae

Description

Empididae is a family of flies with over 3,000 described species occurring worldwide in all the Ecozones but the majority are found in the Holarctic. They are mainly predatory flies like most of their relatives in the Empidoidea, and exhibit a wide range of forms but are generally small to medium-sized, non-metallic and rather bristly.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)9483https://www.gbif.org/species/9483
PaleoBioDB ID (PBDB)139008https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=139008
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Diptera : Empididae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-Holometabola
-Aparaglossata
-Mecopterida
superorderAntliophora
orderDipteraLinnaeus 1758
-NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
-Heterodactyla
-EremoneuraGrimaldi and Cumming 1999
superfamilyEmpidoideaLatreille 1804
familyEmpididaeLatreille 1804
Common Namedagger fly
Scientific NameEmpididae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Empididaefamilybelongs to Dipterastated with evidenceCockerell, 1917
Empididaefamilybelongs to Dipterastated without evidenceCockerell, 1921
Empididaefamilybelongs to Empidoideastated with evidenceKovalev, 1986
Empididaefamilybelongs to Brachycerastated without evidenceCarpenter, 1992
Empididaefamilybelongs to Empidoideastated with evidenceSinclair and Cumming, 2006
Status (PBDB)extant
Taxon Size (PBDB)128
Extant Size (PBDB)27 (21%)
First Recorded Appearance170 - 166 Ma
Middle Jurassic
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)F. M. Carpenter. 1992. Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4:1-655
Common Name(s) Dansflugor, Balloon Flies, Dance Flies, Dance-flies, Dansefluer, Dansefluger, Dansvliegen, Dagger Fly
Wikipediahttps://en.wikipedia.org/wiki/Empididae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Brachystomatinae
subfamily
listed (PBDB)1283.5 Ma
Late/Upper Cretaceous
0 Ma
Extant
Clinocerinae
subfamily
listed (PBDB)256.0 Ma
Paleocene
0 Ma
Extant
Empidinae
subfamily
listed (PBDB)7693.9 Ma
Late/Upper Cretaceous
0 Ma
Extant
Protempidinae
subfamily
listed (PBDB)8170 Ma
Middle Jurassic
125 Ma
Early/Lower Cretaceous
Acallomyia
genus
accepted (GBIF)
listed (PBDB)
137.2 Ma
Eocene
0 Ma
Extant
Achelipoda
genus
accepted (GBIF)No associated record in PBDB
Adipsomyia
genus
accepted (GBIF)No associated record in PBDB
Afroclinocera
genus
accepted (GBIF)No associated record in PBDB
Afrodromia
genus
accepted (GBIF)No associated record in PBDB
Afroempis
genus
accepted (GBIF)No associated record in PBDB
Alavesia
genus
accepted (GBIF)
listed (PBDB)
2105 Ma
Early/Lower Cretaceous
0 Ma
Extant
Allochrotus
genus
accepted (GBIF)No associated record in PBDB
Amictoides
genus
accepted (GBIF)No associated record in PBDB
Anthepiscopus
genus
accepted (GBIF)No associated record in PBDB
Aplomera
genus
accepted (GBIF)No associated record in PBDB
Archiplatypalpus
genus
accepted (GBIF)
listed (PBDB)
185.8 Ma
Late/Upper Cretaceous
85.8 Ma
Late/Upper Cretaceous
Asymphyloptera
genus
accepted (GBIF)No associated record in PBDB
Atelestites
genus
accepted (GBIF)
listed (PBDB)
1129 Ma
Early/Lower Cretaceous
129 Ma
Early/Lower Cretaceous
Atrichopleura
genus
accepted (GBIF)No associated record in PBDB
Avenaphora
genus
accepted (GBIF)
listed (PBDB)
2129 Ma
Early/Lower Cretaceous
85.8 Ma
Late/Upper Cretaceous
Bandella
genus
accepted (GBIF)No associated record in PBDB
Basilorhagio
genus
accepted (GBIF)
listed (PBDB)
1125 Ma
Early/Lower Cretaceous
125 Ma
Early/Lower Cretaceous
Bergenstammia
genus
accepted (GBIF)No associated record in PBDB
Brochella
genus
accepted (GBIF)No associated record in PBDB
Burmitempis
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Chamaedipsia
genus
accepted (GBIF)No associated record in PBDB
Chelifera
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
0 Ma
Extant
Chelipoda
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
0 Ma
Extant
Chelipodozus
genus
accepted (GBIF)No associated record in PBDB
Cladodromia
genus
accepted (GBIF)No associated record in PBDB
Clinocera
genus
accepted (GBIF)No associated record in PBDB
Clinodera
genus
accepted (GBIF)No associated record in PBDB
Clinorhampha
genus
accepted (GBIF)No associated record in PBDB
Colabris
genus
accepted (GBIF)No associated record in PBDB
Coptophlebia
genus
accepted (GBIF)No associated record in PBDB
Cretodromia
genus
accepted (GBIF)
listed (PBDB)
183.5 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Cretomicrophorus
genus
accepted (GBIF)
listed (PBDB)
393.9 Ma
Late/Upper Cretaceous
85.8 Ma
Late/Upper Cretaceous
Cretoplatypalpus
genus
accepted (GBIF)
listed (PBDB)
299.6 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Ctenodrapetis
genus
accepted (GBIF)No associated record in PBDB
Cunomyia
genus
accepted (GBIF)No associated record in PBDB
Cyrtoma
genus
accepted (GBIF)No associated record in PBDB
Deuteragonista
genus
accepted (GBIF)No associated record in PBDB
Dipsomyia
genus
accepted (GBIF)No associated record in PBDB
Dolichocephala
genus
accepted (GBIF)No associated record in PBDB
Doliodromia
genus
accepted (GBIF)No associated record in PBDB
Drapetiella
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Drymodromia
genus
accepted (GBIF)No associated record in PBDB
Dryodromya
genus
accepted (GBIF)No associated record in PBDB
Ecommocydromia
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Edenophorus
genus
accepted (GBIF)No associated record in PBDB
Electrocyrtoma
genus
accepted (GBIF)
listed (PBDB)
199.6 Ma
Late/Upper Cretaceous
99.6 Ma
Late/Upper Cretaceous
Empidadelpha
genus
accepted (GBIF)No associated record in PBDB
Empis
genus
accepted (GBIF)
listed (PBDB)
2493.9 Ma
Late/Upper Cretaceous
0 Ma
Extant
Emplita
genus
accepted (GBIF)
listed (PBDB)
193.9 Ma
Late/Upper Cretaceous
93.9 Ma
Late/Upper Cretaceous
Eugowra
genus
accepted (GBIF)No associated record in PBDB
Euhybos
genus
accepted (GBIF)No associated record in PBDB
Euthyneuriella
genus
accepted (GBIF)
listed (PBDB)
238.0 Ma
Eocene
38.0 Ma
Eocene
Fur
genus
accepted (GBIF)No associated record in PBDB
Gondwanamyia
genus
accepted (GBIF)No associated record in PBDB
Gynatoma
genus
accepted (GBIF)No associated record in PBDB
Hasmona
genus
accepted (GBIF)
listed (PBDB)
1145 Ma
Late Jurassic
145 Ma
Late Jurassic
Hemerodromia
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
0 Ma
Extant
Hesperempis
genus
accepted (GBIF)No associated record in PBDB
Hilara
genus
accepted (GBIF)
listed (PBDB)
638.0 Ma
Eocene
0 Ma
Extant
Hilarempis
genus
accepted (GBIF)No associated record in PBDB
Hilarigona
genus
accepted (GBIF)No associated record in PBDB
Hilarites
genus
accepted (GBIF)
listed (PBDB)
128.1 Ma
Oligocene
28.1 Ma
Oligocene
Holoclera
genus
accepted (GBIF)No associated record in PBDB
Homalocnemimus
genus
accepted (GBIF)
listed (PBDB)
5166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Hormopeza
genus
accepted (GBIF)No associated record in PBDB
Hybomyia
genus
accepted (GBIF)No associated record in PBDB
Hydrodromia
genus
accepted (GBIF)No associated record in PBDB
Hydropeza
genus
accepted (GBIF)No associated record in PBDB
Hypenella
genus
accepted (GBIF)No associated record in PBDB
Hystrichonotus
genus
accepted (GBIF)No associated record in PBDB
Icasma
genus
accepted (GBIF)No associated record in PBDB
Iteaphila
genus
accepted (GBIF)No associated record in PBDB
Kowarzia
genus
accepted (GBIF)No associated record in PBDB
Lamprempis
genus
accepted (GBIF)No associated record in PBDB
Lochmocola
genus
accepted (GBIF)
listed (PBDB)
156.0 Ma
Paleocene
56.0 Ma
Paleocene
Macrostomus
genus
accepted (GBIF)No associated record in PBDB
Meghyperiella
genus
accepted (GBIF)
listed (PBDB)
238.0 Ma
Eocene
38.0 Ma
Eocene
Mesoplatypalpus
genus
accepted (GBIF)
listed (PBDB)
183.5 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Metachela
genus
accepted (GBIF)No associated record in PBDB
Microphorites
genus
accepted (GBIF)
listed (PBDB)
9130 Ma
Early/Lower Cretaceous
55.8 Ma
Eocene
Monodromia
genus
accepted (GBIF)No associated record in PBDB
Munburra
genus
accepted (GBIF)No associated record in PBDB
Nemedromia
genus
accepted (GBIF)
listed (PBDB)
493.9 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Neoplasta
genus
accepted (GBIF)No associated record in PBDB
Neoturonius
genus
accepted (GBIF)
listed (PBDB)
393.9 Ma
Late/Upper Cretaceous
93.9 Ma
Late/Upper Cretaceous
Opeatocerata
genus
accepted (GBIF)No associated record in PBDB
Oreothalia
genus
accepted (GBIF)No associated record in PBDB
Oustaletimyia
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Palaeoleptopeza
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Palaeoparamesia
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Parathalassiella
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Pasitrichotus
genus
accepted (GBIF)No associated record in PBDB
Phaeobalia
genus
accepted (GBIF)No associated record in PBDB
Phaetempis
genus
accepted (GBIF)
listed (PBDB)
2130 Ma
Early/Lower Cretaceous
129 Ma
Early/Lower Cretaceous
Philetus
genus
accepted (GBIF)No associated record in PBDB
Phoneutisca
genus
doubtful (GBIF)
listed (PBDB)
0 Ma
Extant
Phyllodromia
genus
accepted (GBIF)
listed (PBDB)
238.0 Ma
Eocene
0 Ma
Extant
Porphyrochroa
genus
accepted (GBIF)No associated record in PBDB
Proagomyia
genus
accepted (GBIF)No associated record in PBDB
Proclinopyga
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
0 Ma
Extant
Progloma
genus
accepted (GBIF)
listed (PBDB)
137.2 Ma
Eocene
37.2 Ma
Eocene
Prolatomyia
genus
accepted (GBIF)
listed (PBDB)
183.5 Ma
Late/Upper Cretaceous
83.5 Ma
Late/Upper Cretaceous
Protoedalea
genus
accepted (GBIF)
listed (PBDB)
155.8 Ma
Eocene
55.8 Ma
Eocene
Protoreogeton
genus
accepted (GBIF)
listed (PBDB)
1170 Ma
Middle Jurassic
170 Ma
Middle Jurassic
Pseudoacarterus
genus
accepted (GBIF)
listed (PBDB)
193.9 Ma
Late/Upper Cretaceous
93.9 Ma
Late/Upper Cretaceous
Pseudoheleodromia
genus
accepted (GBIF)No associated record in PBDB
Ptilophyllodromia
genus
accepted (GBIF)No associated record in PBDB
Ragas
genus
accepted (GBIF)
listed (PBDB)
838.0 Ma
Eocene
0 Ma
Extant
Retinitus
genus
accepted (GBIF)
listed (PBDB)
185.8 Ma
Late/Upper Cretaceous
85.8 Ma
Late/Upper Cretaceous
Rhamphella
genus
accepted (GBIF)No associated record in PBDB
Rhamphomyia
genus
accepted (GBIF)
listed (PBDB)
3850.3 Ma
Eocene
0 Ma
Extant
Rhyacodromia
genus
accepted (GBIF)No associated record in PBDB
Roederia
genus
accepted (GBIF)No associated record in PBDB
Roederiodes
genus
accepted (GBIF)No associated record in PBDB
Sciodromia
genus
accepted (GBIF)No associated record in PBDB
Sphicosa
genus
accepted (GBIF)No associated record in PBDB
Stuckenbergia
genus
accepted (GBIF)No associated record in PBDB
Tachydroma
genus
accepted (GBIF)No associated record in PBDB
Thinempis
genus
accepted (GBIF)No associated record in PBDB
Thirza
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Trichinites
genus
accepted (GBIF)
listed (PBDB)
1130 Ma
Early/Lower Cretaceous
130 Ma
Early/Lower Cretaceous
Trichoclinocera
genus
accepted (GBIF)No associated record in PBDB
Trichohilara
genus
accepted (GBIF)No associated record in PBDB
Turonempis
genus
accepted (GBIF)
listed (PBDB)
193.9 Ma
Late/Upper Cretaceous
93.9 Ma
Late/Upper Cretaceous
Wiedemania
genus
accepted (GBIF)No associated record in PBDB
Wiedemannia
genus
accepted (GBIF)No associated record in PBDB
Zanclotus
genus
accepted (GBIF)No associated record in PBDB
Bergenitamia multisetosa
species
accepted (GBIF)No associated record in PBDB
Chamedipsia gubernans
species
accepted (GBIF)No associated record in PBDB
Champaedipsia minor
species
accepted (GBIF)No associated record in PBDB
Charadromia arnaudi
species
accepted (GBIF)No associated record in PBDB
Chersdromia megacetes
species
accepted (GBIF)No associated record in PBDB
Chilcottomyia nepalensis
species
accepted (GBIF)No associated record in PBDB
Elaphcopexa upsilon
species
accepted (GBIF)No associated record in PBDB
Hemecodromia brachialis
species
accepted (GBIF)No associated record in PBDB
Hemeredromia raptoria
species
accepted (GBIF)No associated record in PBDB
Hystichonotus revelator
species
accepted (GBIF)No associated record in PBDB
Lundstromiella glaphyra
species
accepted (GBIF)No associated record in PBDB
Oropozella abdominalis
species
accepted (GBIF)No associated record in PBDB
Oropozella annulitarsis
species
accepted (GBIF)No associated record in PBDB
Oropozella antennata
species
accepted (GBIF)No associated record in PBDB
Oropozella arara
species
accepted (GBIF)No associated record in PBDB
Oropozella bifurcata
species
accepted (GBIF)No associated record in PBDB
Oropozella biloba
species
accepted (GBIF)No associated record in PBDB
Oropozella diminuloruma
species
accepted (GBIF)No associated record in PBDB
Oropozella loripes
species
accepted (GBIF)No associated record in PBDB
Oropozella marginata
species
accepted (GBIF)No associated record in PBDB
Oropozella metallica
species
accepted (GBIF)No associated record in PBDB
Oropozella tanycera
species
accepted (GBIF)No associated record in PBDB
Oropozella taquara
species
accepted (GBIF)No associated record in PBDB
Oropozella tricuspicata
species
accepted (GBIF)No associated record in PBDB
Phyletus schizophorus
species
accepted (GBIF)No associated record in PBDB
Tachyeurpis cinerea
species
accepted (GBIF)No associated record in PBDB

Obsolete Names

NameSourceTaxon RankTaxonomy
Empedidae GBIFfamilyAnimalia : Arthropoda : Insecta : Diptera : Empedidae
Tachydromia stilpon Meunier, 1899GBIFspeciesAnimalia : Arthropoda : Insecta : Diptera : Empididae : Tachydromia stilpon

Synonymy List

YearName and Author
1804Empididae Latreille p. 310
1899Tachydromia stilpon Meunier p. 178
1908Brachystomatidae Melander
1917Empedidae Cockerell p. 367
1921Empididae Cockerell
1986Empididae Kovalev p. 152
YearName and Author
1992Empididae Carpenter
2006Empididae Sinclair and Cumming
2006Brachystomatidae Sinclair and Cumming p. 80
2011Brachystomatidae Sinclair et al.
2011Empididae Wiegmann et al.
2018Empididae Wahlberg and Johanson

References

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655
Meunier F. Révision des diptères fossiles types de Loew conservés au Musée Provincial de Koenigsberg, Miscellanea Entomologica 7, 169-182
Cockerell T. D. A. (1917) Arthropods in Burmese amber, American Journal of Science, Series 4 44 263, 360-368
Cockerell T. D. A. (1921) Eocene Insects from the Rocky Mountains, Proceedings of the United States National Museum 57, 233-260
Kovalev V. G. (1986) Bibionomorpha and Asilomorpha, in Nasekomye v rannemelovykh ekosistemakh zapadnoy Mongolii, The Joint Soviet-Mongolian Palaeontological Expedition 28, 125-154
Sinclair B. J., Cumming J. M. (2006) The morphology, higher-level phylogeny and classification of the Empidoidea (Diptera), Zootaxa 1180, 1-172
Sinclair B. J., Brooks S. E., et al (2011) Revision of the Nearctic species of Heleodromia (Diptera: Empidoidea: Brachystomatidae), The Canadian Entomologist 143, 629-651
Wiegmann B. M., Trautwein M. D., et al (2011) Episodic radiations in the fly tree of life, Proceedings of the National Academy of Sciences 108, 5690-5695
Wahlberg E., Johanson K. A. (2018) Molecular phylogenetics reveals novel relationships within Empidoidea (Diptera), Systematic Entomology 43, 619-636
Beuk, P.L.Th. (ed.) (2002) - via Taxa Watermanagement the Netherlands (TWN)
Smith, K.G.V. (1989) - via Taxa Watermanagement the Netherlands (TWN)
Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas
Sinclair, J.M. (2006) The morphology, higher-level phylogeny and classificationof the Empidoidea (Diptera). Zootaxa 1180, 172 pp. Magnolia Press, Auckland - via Dyntaxa. Svensk taxonomisk databas
De Pauw, N. & Vannevel, R. (eds.) (1991) - via Taxa Watermanagement the Netherlands (TWN)
Nilsson, A. (ed.) (1997) - via Taxa Watermanagement the Netherlands (TWN)
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
some verified marine genera (AJR, Nov 2010) - via The Interim Register of Marine and Nonmarine Genera
Grootaert, P.; De Bruyn, L.; De Meyer, M. (1991). Catalogue of the Diptera of Belgium. Studiedocumenten van het Koninklijk Belgisch Instituut voor Natuurwetenschappen. 70: 338pp. - via World Register of Marine Species
Vockeroth, J. R.,Peterson, B. V.,Knutson, L. V.,Teskey, H. J.,McAlpine, J. F.,Steyskal, G. C.,Shewell, G. E.,Wood, D. M. (1981) Chapter 47. Empididae - via The Diptera Site
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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