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Pterostichus

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Description

Pterostichus is a very large genus of ground beetles with a Holarctic distribution in the subfamily Harpalinae. It has over 1,100 species. The beetles are predatory, but sometimes feed on strawberries. They can be found under rocks and prefer slightly moist, sandy soil although preferences differ between species. Pterostichus contains the following species:

From Wikipedia article at https://en.wikipedia.org/wiki/Pterostichus, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	7367006	https://www.gbif.org/species/7367006

Rank

genus

Taxonomy (GBIF)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Pterostichus

Taxonomic Status (GBIF)

accepted

Classification
(GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
genus	Pterostichus	Bonelli 1810

Scientific Name

Pterostichus Bonelli, 1810

Name Published In

Bonelli, Franco A. 1810. Observations entomologiques. Première partie (cicindélètes et portion des carabiques). Félix Galletti, Turin.: 1-58; Tabula synoptica.

Wikipedia

https://en.wikipedia.org/wiki/Pterostichus

External Images

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Pterostichus aanistschenkoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus aapsorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus abaciformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus abagonensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus abasarukini species	accepted (GBIF)	No associated record in PBDB
Pterostichus abasinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus abaxoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus abensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus abishirensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus abnormis species	accepted (GBIF) listed (PBDB)		3	2.59 Ma Pleistocene	0 Ma Extant
Pterostichus abrogatus ✝ species	accepted (GBIF) listed (PBDB)		3	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus abudarini species	accepted (GBIF)	No associated record in PBDB
Pterostichus acrogonus species	accepted (GBIF)	No associated record in PBDB
Pterostichus acuspina species	accepted (GBIF)	No associated record in PBDB
Pterostichus acutidens species	accepted (GBIF)	No associated record in PBDB
Pterostichus acutipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus adatarasanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ademidovae species	accepted (GBIF)	No associated record in PBDB
Pterostichus adoxus species	accepted (GBIF)	No associated record in PBDB
Pterostichus adstrictus species	accepted (GBIF)	No associated record in PBDB
Pterostichus adulterinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus aemiliae species	accepted (GBIF)	No associated record in PBDB
Pterostichus aeneocupreus species	accepted (GBIF)	No associated record in PBDB
Pterostichus aenescens species	accepted (GBIF)	No associated record in PBDB
Pterostichus aereipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus aethiops species	accepted (GBIF)	No associated record in PBDB
Pterostichus agilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus agonus species	accepted (GBIF)	No associated record in PBDB
Pterostichus aibgensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus aimaki species	accepted (GBIF)	No associated record in PBDB
Pterostichus ainus species	accepted (GBIF)	No associated record in PBDB
Pterostichus akitai species	accepted (GBIF)	No associated record in PBDB
Pterostichus akkusianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus akozyrevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus aksekianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus alacer species	accepted (GBIF)	No associated record in PBDB
Pterostichus alexandrovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus alexeji species	accepted (GBIF)	No associated record in PBDB
Pterostichus algidus species	accepted (GBIF)	No associated record in PBDB
Pterostichus altaianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus altaicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus altaiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus amadeus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amagisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amanoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus amaroides species	accepted (GBIF)	No associated record in PBDB
Pterostichus ambigenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ambiguus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amblypterus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amethystinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amoenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus amurensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus analis species	accepted (GBIF)	No associated record in PBDB
Pterostichus anatolicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus andreae species	accepted (GBIF)	No associated record in PBDB
Pterostichus andreinii species	accepted (GBIF)	No associated record in PBDB
Pterostichus andrewesi species	accepted (GBIF)	No associated record in PBDB
Pterostichus angoarnigi species	accepted (GBIF)	No associated record in PBDB
Pterostichus angustus species	accepted (GBIF)	No associated record in PBDB
Pterostichus annamita species	accepted (GBIF)	No associated record in PBDB
Pterostichus annosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus anomostriatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus antepunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus anthracinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus apenninus species	accepted (GBIF)	No associated record in PBDB
Pterostichus apfelbecki species	accepted (GBIF)	No associated record in PBDB
Pterostichus apiculatiphallus species	accepted (GBIF)	No associated record in PBDB
Pterostichus aralarensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus araraticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus arator species	accepted (GBIF)	No associated record in PBDB
Pterostichus arcanoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus arcanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus arcticola species	accepted (GBIF)	No associated record in PBDB
Pterostichus arcuaticarinatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus argutoriformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus aristochroides species	accepted (GBIF)	No associated record in PBDB
Pterostichus armenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus arnoldii species	accepted (GBIF)	No associated record in PBDB
Pterostichus arribasi species	accepted (GBIF)	No associated record in PBDB
Pterostichus arrowi species	accepted (GBIF)	No associated record in PBDB
Pterostichus arrowianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus arsenjevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus asahinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus asiaticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus asturicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus astutus species	accepted (GBIF)	No associated record in PBDB
Pterostichus asymmetricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus aterrimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus atrox species	accepted (GBIF)	No associated record in PBDB
Pterostichus aubryi species	accepted (GBIF)	No associated record in PBDB
Pterostichus audax species	accepted (GBIF)	No associated record in PBDB
Pterostichus auratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus auriga species	accepted (GBIF)	No associated record in PBDB
Pterostichus avaricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus baenningeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus balachowskyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus baldwini species	accepted (GBIF)	No associated record in PBDB
Pterostichus balikliensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus balthasari species	accepted (GBIF)	No associated record in PBDB
Pterostichus balu species	accepted (GBIF)	No associated record in PBDB
Pterostichus bandotaro species	accepted (GBIF)	No associated record in PBDB
Pterostichus barbarae species	accepted (GBIF)	No associated record in PBDB
Pterostichus barbarinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus barlensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus barri species	accepted (GBIF)	No associated record in PBDB
Pterostichus barryorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus basilobatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus basipunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus batesi species	accepted (GBIF)	No associated record in PBDB
Pterostichus belizini species	accepted (GBIF)	No associated record in PBDB
Pterostichus bellatrix species	accepted (GBIF)	No associated record in PBDB
Pterostichus belousovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus beneshi species	accepted (GBIF)	No associated record in PBDB
Pterostichus berezowskii species	accepted (GBIF)	No associated record in PBDB
Pterostichus besucheti species	accepted (GBIF)	No associated record in PBDB
Pterostichus beyeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus bhadravati species	accepted (GBIF)	No associated record in PBDB
Pterostichus bhunetansis species	accepted (GBIF)	No associated record in PBDB
Pterostichus bhutanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus bicolor species	accepted (GBIF)	No associated record in PBDB
Pterostichus bielzii species	accepted (GBIF)	No associated record in PBDB
Pterostichus biexcisus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bifidiphallus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bifoveolatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus biocryus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bischoffianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bisetosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bisulcitarsis species	accepted (GBIF)	No associated record in PBDB
Pterostichus bituberculatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus blandulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus blodgettensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus blumenthali species	accepted (GBIF)	No associated record in PBDB
Pterostichus borcka species	accepted (GBIF)	No associated record in PBDB
Pterostichus bousqueti species	accepted (GBIF)	No associated record in PBDB
Pterostichus bowanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus brachylobus species	accepted (GBIF)	No associated record in PBDB
Pterostichus brancuccii species	accepted (GBIF)	No associated record in PBDB
Pterostichus brevicornis species	accepted (GBIF)	No associated record in PBDB
Pterostichus brevilama species	accepted (GBIF)	No associated record in PBDB
Pterostichus brevipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus brevis species	accepted (GBIF)	No associated record in PBDB
Pterostichus breviusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus brittoni species	accepted (GBIF)	No associated record in PBDB
Pterostichus bruckii species	accepted (GBIF)	No associated record in PBDB
Pterostichus brunneipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus bryanti species	accepted (GBIF)	No associated record in PBDB
Pterostichus bryantoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus buglaniensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus bullatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus bungei species	accepted (GBIF)	No associated record in PBDB
Pterostichus bureli species	accepted (GBIF)	No associated record in PBDB
Pterostichus burkhan species	accepted (GBIF)	No associated record in PBDB
Pterostichus burmeisteri species	accepted (GBIF)	No associated record in PBDB
Pterostichus bythiniensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cacumensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus californicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus caligans species	accepted (GBIF)	No associated record in PBDB
Pterostichus calvitarsis species	accepted (GBIF)	No associated record in PBDB
Pterostichus campbelli species	accepted (GBIF)	No associated record in PBDB
Pterostichus canallatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cancellatus species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0 Ma Extant
Pterostichus cantabricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cantalicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus capitatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus capito species	accepted (GBIF)	No associated record in PBDB
Pterostichus capitolinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus caribou species	accepted (GBIF)	No associated record in PBDB
Pterostichus carinatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus carolinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus carri species	accepted (GBIF)	No associated record in PBDB
Pterostichus carsticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus casalei species	accepted (GBIF)	No associated record in PBDB
Pterostichus casaleianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus caspius species	accepted (GBIF)	No associated record in PBDB
Pterostichus castaneus species	accepted (GBIF)	No associated record in PBDB
Pterostichus castanipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus castor species	accepted (GBIF)	No associated record in PBDB
Pterostichus catalonicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus catei species	accepted (GBIF)	No associated record in PBDB
Pterostichus cathaicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus caucasicola species	accepted (GBIF)	No associated record in PBDB
Pterostichus caucasicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus caudicalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cavazzutii species	accepted (GBIF)	No associated record in PBDB
Pterostichus cavicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cecchiniae species	accepted (GBIF)	No associated record in PBDB
Pterostichus cervenkai species	accepted (GBIF)	No associated record in PBDB
Pterostichus chainapaani species	accepted (GBIF)	No associated record in PBDB
Pterostichus chameleon species	accepted (GBIF)	No associated record in PBDB
Pterostichus champenoisi species	accepted (GBIF)	No associated record in PBDB
Pterostichus championi species	accepted (GBIF)	No associated record in PBDB
Pterostichus chasautianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chechcirensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus chefsuricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chenpengi species	accepted (GBIF)	No associated record in PBDB
Pterostichus chinkiangensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus chinlingshanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chipewyan species	accepted (GBIF)	No associated record in PBDB
Pterostichus chogyesanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chokaisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chotanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus chujoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus chujoiellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus chungkingi species	accepted (GBIF)	No associated record in PBDB
Pterostichus chydaeus species	accepted (GBIF)	No associated record in PBDB
Pterostichus circulosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus clepsydra species	accepted (GBIF)	No associated record in PBDB
Pterostichus cochlearis species	accepted (GBIF)	No associated record in PBDB
Pterostichus colasi species	accepted (GBIF)	No associated record in PBDB
Pterostichus colchicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus colonus species	accepted (GBIF)	No associated record in PBDB
Pterostichus comatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cometes species	accepted (GBIF)	No associated record in PBDB
Pterostichus commixtiformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus commutabilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus comorus species	accepted (GBIF)	No associated record in PBDB
Pterostichus compressus species	accepted (GBIF)	No associated record in PBDB
Pterostichus conaensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus confucius species	accepted (GBIF)	No associated record in PBDB
Pterostichus congestus species	accepted (GBIF)	No associated record in PBDB
Pterostichus consanguineus species	accepted (GBIF)	No associated record in PBDB
Pterostichus consors species	accepted (GBIF)	No associated record in PBDB
Pterostichus coracinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cordaticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cordatissimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cordifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus coreicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus corrusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus coruscus species	accepted (GBIF)	No associated record in PBDB
Pterostichus corvinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus costatus species	accepted (GBIF) listed (PBDB)		4	11.6 Ma Miocene	0 Ma Extant
Pterostichus crassiapex species	accepted (GBIF)	No associated record in PBDB
Pterostichus crassiusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus craterensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cratocephalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus crenicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus crenulatopunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus creper species	accepted (GBIF)	No associated record in PBDB
Pterostichus cribratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cristatoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus cristatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cristicaudis species	accepted (GBIF)	No associated record in PBDB
Pterostichus cryobioides species	accepted (GBIF)	No associated record in PBDB
Pterostichus cucujinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cunibakus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cursor species	accepted (GBIF)	No associated record in PBDB
Pterostichus curtatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus curvatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cylindricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus cylindriformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus daghestanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus daihizanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus daisenicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus dandongensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus dandonis species	accepted (GBIF)	No associated record in PBDB
Pterostichus darvazicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus datshenkoae species	accepted (GBIF)	No associated record in PBDB
Pterostichus dauricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus davidi species	accepted (GBIF)	No associated record in PBDB
Pterostichus decastriensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus deceptrix species	accepted (GBIF)	No associated record in PBDB
Pterostichus defossus species	accepted (GBIF)	No associated record in PBDB
Pterostichus deino species	accepted (GBIF)	No associated record in PBDB
Pterostichus dentellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus denticaudis species	accepted (GBIF)	No associated record in PBDB
Pterostichus dentifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus depilatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus depletus ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus depressidorsis species	accepted (GBIF)	No associated record in PBDB
Pterostichus destitutus ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus destructus ✝ species	accepted (GBIF) listed (PBDB)		2	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus deuvei species	accepted (GBIF)	No associated record in PBDB
Pterostichus deuvesianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus devillei species	accepted (GBIF)	No associated record in PBDB
Pterostichus dhorpatanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus diana species	accepted (GBIF)	No associated record in PBDB
Pterostichus diligendus species	accepted (GBIF)	No associated record in PBDB
Pterostichus diligens species	accepted (GBIF)	No associated record in PBDB
Pterostichus dilutipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus dipojranus species	accepted (GBIF)	No associated record in PBDB
Pterostichus discrepans species	accepted (GBIF)	No associated record in PBDB
Pterostichus dissimilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus distinctissimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus distinctus ✝ species	accepted (GBIF) listed (PBDB)		1	28.4 Ma Oligocene	28.4 Ma Oligocene
Pterostichus diversus species	accepted (GBIF)	No associated record in PBDB
Pterostichus dolens species	accepted (GBIF)	No associated record in PBDB
Pterostichus doris species	accepted (GBIF)	No associated record in PBDB
Pterostichus dorjulensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus dormitans ✝ species	accepted (GBIF) listed (PBDB)		2	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus dostali species	accepted (GBIF)	No associated record in PBDB
Pterostichus drescheri species	accepted (GBIF)	No associated record in PBDB
Pterostichus drescoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus dudkoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus dufourii species	accepted (GBIF)	No associated record in PBDB
Pterostichus dulcis species	accepted (GBIF)	No associated record in PBDB
Pterostichus dundai species	accepted (GBIF)	No associated record in PBDB
Pterostichus duratii species	accepted (GBIF)	No associated record in PBDB
Pterostichus dux species	accepted (GBIF)	No associated record in PBDB
Pterostichus ebeninus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ebenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus eboshiyamanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ecarinatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ehlersi species	accepted (GBIF)	No associated record in PBDB
Pterostichus elmbergi species	accepted (GBIF)	No associated record in PBDB
Pterostichus elongatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus emei species	accepted (GBIF)	No associated record in PBDB
Pterostichus empetricola species	accepted (GBIF)	No associated record in PBDB
Pterostichus enasanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus enodis species	accepted (GBIF)	No associated record in PBDB
Pterostichus enyo species	accepted (GBIF)	No associated record in PBDB
Pterostichus eobius species	accepted (GBIF)	No associated record in PBDB
Pterostichus eoyoritomus species	accepted (GBIF)	No associated record in PBDB
Pterostichus epiroticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus eriwanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus esakii species	accepted (GBIF)	No associated record in PBDB
Pterostichus eschscholtzii species	accepted (GBIF)	No associated record in PBDB
Pterostichus espagnoli species	accepted (GBIF)	No associated record in PBDB
Pterostichus espanoli species	accepted (GBIF)	No associated record in PBDB
Pterostichus esuriens species	accepted (GBIF)	No associated record in PBDB
Pterostichus euxinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus exceptus species	accepted (GBIF)	No associated record in PBDB
Pterostichus exilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus eximius species	accepted (GBIF)	No associated record in PBDB
Pterostichus exochus species	accepted (GBIF)	No associated record in PBDB
Pterostichus expeditus species	accepted (GBIF)	No associated record in PBDB
Pterostichus externepunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus falcispinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus faldermanni species	accepted (GBIF)	No associated record in PBDB
Pterostichus fallettii species	accepted (GBIF)	No associated record in PBDB
Pterostichus falli species	accepted (GBIF)	No associated record in PBDB
Pterostichus farkaci species	accepted (GBIF)	No associated record in PBDB
Pterostichus fasciatopunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus faunus species	accepted (GBIF)	No associated record in PBDB
Pterostichus femoralis species	accepted (GBIF)	No associated record in PBDB
Pterostichus fencli species	accepted (GBIF)	No associated record in PBDB
Pterostichus fenestratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus fenyesi species	accepted (GBIF)	No associated record in PBDB
Pterostichus ferghanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus fernquisti ✝ species	accepted (GBIF) listed (PBDB)		1	16.0 Ma Miocene	16.0 Ma Miocene
Pterostichus ferreri species	accepted (GBIF)	No associated record in PBDB
Pterostichus ferreroi species	accepted (GBIF)	No associated record in PBDB
Pterostichus filipjevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus filum species	accepted (GBIF)	No associated record in PBDB
Pterostichus findelii species	accepted (GBIF)	No associated record in PBDB
Pterostichus firmus species	accepted (GBIF)	No associated record in PBDB
Pterostichus flavofemoratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus focarilei species	accepted (GBIF)	No associated record in PBDB
Pterostichus fornicatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus foveolatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus fractus ✝ species	accepted (GBIF) listed (PBDB)		2	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Pterostichus franzi species	accepted (GBIF)	No associated record in PBDB
Pterostichus fritzhiekei species	accepted (GBIF)	No associated record in PBDB
Pterostichus fuchsi species	accepted (GBIF)	No associated record in PBDB
Pterostichus fujimurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus fujisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus fujitai species	accepted (GBIF)	No associated record in PBDB
Pterostichus fukube species	accepted (GBIF)	No associated record in PBDB
Pterostichus fuligineus species	accepted (GBIF)	No associated record in PBDB
Pterostichus fulvescens species	accepted (GBIF)	No associated record in PBDB
Pterostichus funakoshii species	accepted (GBIF)	No associated record in PBDB
Pterostichus funestes species	accepted (GBIF)	No associated record in PBDB
Pterostichus fuscicornis species	accepted (GBIF)	No associated record in PBDB
Pterostichus gagates species	accepted (GBIF)	No associated record in PBDB
Pterostichus galae species	accepted (GBIF)	No associated record in PBDB
Pterostichus galaecianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus galinae species	accepted (GBIF)	No associated record in PBDB
Pterostichus gallega species	accepted (GBIF)	No associated record in PBDB
Pterostichus gallopavo species	accepted (GBIF)	No associated record in PBDB
Pterostichus ganesh species	accepted (GBIF)	No associated record in PBDB
Pterostichus ganja species	accepted (GBIF)	No associated record in PBDB
Pterostichus gaoligongensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus gassanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus geberti species	accepted (GBIF)	No associated record in PBDB
Pterostichus geojensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus georgi species	accepted (GBIF)	No associated record in PBDB
Pterostichus gerdi species	accepted (GBIF)	No associated record in PBDB
Pterostichus gerstlensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ghilarovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus ghilianii species	accepted (GBIF)	No associated record in PBDB
Pterostichus giacomazzoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus gibbicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus giresuni species	accepted (GBIF)	No associated record in PBDB
Pterostichus glabricollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus glabripennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus glacialis species	accepted (GBIF)	No associated record in PBDB
Pterostichus glaferi species	accepted (GBIF)	No associated record in PBDB
Pterostichus gliscans species	accepted (GBIF)	No associated record in PBDB
Pterostichus glukhomanka species	accepted (GBIF)	No associated record in PBDB
Pterostichus gobettii species	accepted (GBIF)	No associated record in PBDB
Pterostichus gompanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus gongga species	accepted (GBIF)	No associated record in PBDB
Pterostichus goriensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus goschi species	accepted (GBIF)	No associated record in PBDB
Pterostichus gotoensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus gracilior species	accepted (GBIF)	No associated record in PBDB
Pterostichus gracilis species	accepted (GBIF) listed (PBDB)		3	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus grajus species	accepted (GBIF)	No associated record in PBDB
Pterostichus gravis species	accepted (GBIF)	No associated record in PBDB
Pterostichus gromykoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus guizhouensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus gurnetensis ✝ species	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Pterostichus gwervaenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus gyrosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus haesitatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus haftorni species	accepted (GBIF)	No associated record in PBDB
Pterostichus hagenbachii species	accepted (GBIF)	No associated record in PBDB
Pterostichus hakusanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus haptoderoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus harmandi species	accepted (GBIF)	No associated record in PBDB
Pterostichus harponifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus hartmanni species	accepted (GBIF)	No associated record in PBDB
Pterostichus hatchi species	accepted (GBIF)	No associated record in PBDB
Pterostichus hedkeinekus species	accepted (GBIF)	No associated record in PBDB
Pterostichus heilongjiangensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus heinzi species	accepted (GBIF)	No associated record in PBDB
Pterostichus heinzianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus hejkali species	accepted (GBIF)	No associated record in PBDB
Pterostichus heptapotamicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus herculaneus species	accepted (GBIF)	No associated record in PBDB
Pterostichus herzi species	accepted (GBIF)	No associated record in PBDB
Pterostichus hidanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus hikosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus himifuriho species	accepted (GBIF)	No associated record in PBDB
Pterostichus hiramatsui species	accepted (GBIF)	No associated record in PBDB
Pterostichus hirasawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus hirobane species	accepted (GBIF)	No associated record in PBDB
Pterostichus hisamatsui species	accepted (GBIF)	No associated record in PBDB
Pterostichus hoelli species	accepted (GBIF)	No associated record in PBDB
Pterostichus homalonotus species	accepted (GBIF)	No associated record in PBDB
Pterostichus honnoratii species	accepted (GBIF)	No associated record in PBDB
Pterostichus hoplites species	accepted (GBIF)	No associated record in PBDB
Pterostichus hornii species	accepted (GBIF)	No associated record in PBDB
Pterostichus horstweiperti species	accepted (GBIF)	No associated record in PBDB
Pterostichus horvatovichi species	accepted (GBIF)	No associated record in PBDB
Pterostichus hozumii species	accepted (GBIF)	No associated record in PBDB
Pterostichus huashanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus hubeicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus hudsonicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus humidulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus humilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus hungaricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus hypogeus species	accepted (GBIF)	No associated record in PBDB
Pterostichus iberlovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus idahoae species	accepted (GBIF)	No associated record in PBDB
Pterostichus ienishteai species	accepted (GBIF)	No associated record in PBDB
Pterostichus ignavus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ikizderensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ikukoae species	accepted (GBIF)	No associated record in PBDB
Pterostichus illigeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus illustris species	accepted (GBIF)	No associated record in PBDB
Pterostichus imasakai species	accepted (GBIF)	No associated record in PBDB
Pterostichus imitatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus immarginatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus impressicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus impressus species	accepted (GBIF)	No associated record in PBDB
Pterostichus imurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus inanis species	accepted (GBIF)	No associated record in PBDB
Pterostichus incommodus species	accepted (GBIF)	No associated record in PBDB
Pterostichus inermis species	accepted (GBIF)	No associated record in PBDB
Pterostichus infernalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus infimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ingusha species	accepted (GBIF)	No associated record in PBDB
Pterostichus innshanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus inopinatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus inopinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus insidatrix species	accepted (GBIF)	No associated record in PBDB
Pterostichus insidiator species	accepted (GBIF)	No associated record in PBDB
Pterostichus insignicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus interruptus species	accepted (GBIF)	No associated record in PBDB
Pterostichus irideus species	accepted (GBIF)	No associated record in PBDB
Pterostichus iripennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus isabellae species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishiharai species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishii species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishikawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishikawaioides species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishizuchiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ishizukai species	accepted (GBIF)	No associated record in PBDB
Pterostichus isolatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus isumiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus itoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus iwakiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ixion species	accepted (GBIF)	No associated record in PBDB
Pterostichus jacobinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jacobsoni species	accepted (GBIF)	No associated record in PBDB
Pterostichus jaechi species	accepted (GBIF)	No associated record in PBDB
Pterostichus jaechianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jakobsonianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jaljaleensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus janatai species	accepted (GBIF)	No associated record in PBDB
Pterostichus janbritoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus jani species	accepted (GBIF)	No associated record in PBDB
Pterostichus jankowskyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus janoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus japonicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jelepus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jiricola species	accepted (GBIF)	No associated record in PBDB
Pterostichus jogaesanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus johnsoni species	accepted (GBIF)	No associated record in PBDB
Pterostichus josephi species	accepted (GBIF)	No associated record in PBDB
Pterostichus juga species	accepted (GBIF)	No associated record in PBDB
Pterostichus jugicola species	accepted (GBIF)	No associated record in PBDB
Pterostichus jugivagus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jungens species	accepted (GBIF)	No associated record in PBDB
Pterostichus jureceki species	accepted (GBIF)	No associated record in PBDB
Pterostichus jurecekianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus jurinei species	accepted (GBIF)	No associated record in PBDB
Pterostichus justusii species	accepted (GBIF)	No associated record in PBDB
Pterostichus kadjaranze species	accepted (GBIF)	No associated record in PBDB
Pterostichus kadleci species	accepted (GBIF)	No associated record in PBDB
Pterostichus kadoudali species	accepted (GBIF)	No associated record in PBDB
Pterostichus kajimurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kalabi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kalhys species	accepted (GBIF)	No associated record in PBDB
Pterostichus kalo species	accepted (GBIF)	No associated record in PBDB
Pterostichus kambaiti species	accepted (GBIF)	No associated record in PBDB
Pterostichus kamtschaticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kangchenjunga species	accepted (GBIF)	No associated record in PBDB
Pterostichus kaniei species	accepted (GBIF)	No associated record in PBDB
Pterostichus kaninensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus karasawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus karateghinicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kataevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus katashinensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus kaynashensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus keltiekus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kerzhneri species	accepted (GBIF)	No associated record in PBDB
Pterostichus kiangsu species	accepted (GBIF)	No associated record in PBDB
Pterostichus kiiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus kimurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kiritshenkoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kirschenblatti species	accepted (GBIF)	No associated record in PBDB
Pterostichus kitakamisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus klapperichi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kleinfeldi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kleinfeldianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus koenigi species	accepted (GBIF)	No associated record in PBDB
Pterostichus koenigianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus koheii species	accepted (GBIF)	No associated record in PBDB
Pterostichus kokeilii species	accepted (GBIF)	No associated record in PBDB
Pterostichus kolosovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kolymensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus komalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus komiyai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kongosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kopetzi species	accepted (GBIF)	No associated record in PBDB
Pterostichus korgei species	accepted (GBIF)	No associated record in PBDB
Pterostichus korolkowi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kosakai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kotzebuei species	accepted (GBIF)	No associated record in PBDB
Pterostichus krali species	accepted (GBIF)	No associated record in PBDB
Pterostichus krasnopolensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus krulikovskyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kucerai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kultianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kurentzovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus kurikomasanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus kurosai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kurosawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus kuwolsanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus kvirensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus laborans species	accepted (GBIF)	No associated record in PBDB
Pterostichus labzuki species	accepted (GBIF)	No associated record in PBDB
Pterostichus lacertus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lachrymosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lacunosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus laevigatus ✝ species	accepted (GBIF) listed (PBDB)		1	1.80 Ma Pleistocene	1.80 Ma Pleistocene
Pterostichus laevipunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus laferi species	accepted (GBIF)	No associated record in PBDB
Pterostichus lama species	accepted (GBIF)	No associated record in PBDB
Pterostichus lamuticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lanei species	accepted (GBIF)	No associated record in PBDB
Pterostichus lanista species	accepted (GBIF)	No associated record in PBDB
Pterostichus larisae species	accepted (GBIF)	No associated record in PBDB
Pterostichus lasserrei species	accepted (GBIF)	No associated record in PBDB
Pterostichus lassulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus latemarginatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus laticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus latifianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus latistylis species	accepted (GBIF)	No associated record in PBDB
Pterostichus latitemporis species	accepted (GBIF)	No associated record in PBDB
Pterostichus latiusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lattini species	accepted (GBIF)	No associated record in PBDB
Pterostichus ledenikensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus leica species	accepted (GBIF)	No associated record in PBDB
Pterostichus leonisi species	accepted (GBIF)	No associated record in PBDB
Pterostichus leptis species	accepted (GBIF)	No associated record in PBDB
Pterostichus lesticoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus letensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus levadensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus leviculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus licenti species	accepted (GBIF)	No associated record in PBDB
Pterostichus liciniformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus lidarus species	accepted (GBIF)	No associated record in PBDB
Pterostichus liguricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lineatopunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lingshanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus liodactylus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lobatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lodosi species	accepted (GBIF)	No associated record in PBDB
Pterostichus lolo species	accepted (GBIF)	No associated record in PBDB
Pterostichus lombardus species	accepted (GBIF)	No associated record in PBDB
Pterostichus longinquus species	accepted (GBIF)	No associated record in PBDB
Pterostichus longipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus longipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus longiusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lubricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lucidus species	accepted (GBIF)	No associated record in PBDB
Pterostichus luctuosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus lugi species	accepted (GBIF)	No associated record in PBDB
Pterostichus lukjanovitshi species	accepted (GBIF)	No associated record in PBDB
Pterostichus lumensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus lustrans species	accepted (GBIF)	No associated record in PBDB
Pterostichus lutschniki species	accepted (GBIF)	No associated record in PBDB
Pterostichus lutshnikianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus macedonicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus macer species	accepted (GBIF)	No associated record in PBDB
Pterostichus machulkai species	accepted (GBIF)	No associated record in PBDB
Pterostichus macrocephalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus macrogenys species	accepted (GBIF)	No associated record in PBDB
Pterostichus macrothorax species	accepted (GBIF)	No associated record in PBDB
Pterostichus maderi species	accepted (GBIF)	No associated record in PBDB
Pterostichus madidus species	accepted (GBIF)	No associated record in PBDB
Pterostichus maeklini species	accepted (GBIF)	No associated record in PBDB
Pterostichus magoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus magus species	accepted (GBIF)	No associated record in PBDB
Pterostichus maichensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus makalu species	accepted (GBIF)	No associated record in PBDB
Pterostichus makolskii species	accepted (GBIF)	No associated record in PBDB
Pterostichus malaise species	accepted (GBIF)	No associated record in PBDB
Pterostichus malaisei species	accepted (GBIF)	No associated record in PBDB
Pterostichus malissorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus malkini species	accepted (GBIF)	No associated record in PBDB
Pterostichus mancus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mandibularoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus mandzhuricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mannerheimii species	accepted (GBIF)	No associated record in PBDB
Pterostichus marani species	accepted (GBIF)	No associated record in PBDB
Pterostichus marginellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mariae species	accepted (GBIF)	No associated record in PBDB
Pterostichus mariamae species	accepted (GBIF)	No associated record in PBDB
Pterostichus marinensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus masahiroi species	accepted (GBIF)	No associated record in PBDB
Pterostichus masatakai species	accepted (GBIF)	No associated record in PBDB
Pterostichus mascareauxi species	accepted (GBIF)	No associated record in PBDB
Pterostichus mashidai species	accepted (GBIF)	No associated record in PBDB
Pterostichus masidai species	accepted (GBIF)	No associated record in PBDB
Pterostichus masumotoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus matchai species	accepted (GBIF)	No associated record in PBDB
Pterostichus matsumurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus mattolenis species	accepted (GBIF)	No associated record in PBDB
Pterostichus mattolensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus matveichuki species	accepted (GBIF)	No associated record in PBDB
Pterostichus maurusiacus species	accepted (GBIF)	No associated record in PBDB
Pterostichus maximus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mediocris species	accepted (GBIF)	No associated record in PBDB
Pterostichus medvedevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus megacephalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus megaloderus species	accepted (GBIF)	No associated record in PBDB
Pterostichus meisteri species	accepted (GBIF)	No associated record in PBDB
Pterostichus melanarius species	accepted (GBIF) listed (PBDB)		3	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus melanodes species	accepted (GBIF)	No associated record in PBDB
Pterostichus melas species	accepted (GBIF)	No associated record in PBDB
Pterostichus mellyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus menetriesii species	accepted (GBIF)	No associated record in PBDB
Pterostichus mercedianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus merkli species	accepted (GBIF)	No associated record in PBDB
Pterostichus meskheticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mewakholensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus meyeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus micans species	accepted (GBIF)	No associated record in PBDB
Pterostichus michailovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus microbus species	accepted (GBIF)	No associated record in PBDB
Pterostichus microcephalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus microphthalmus species	accepted (GBIF)	No associated record in PBDB
Pterostichus microps species	accepted (GBIF)	No associated record in PBDB
Pterostichus middendorffi species	accepted (GBIF)	No associated record in PBDB
Pterostichus miehei species	accepted (GBIF)	No associated record in PBDB
Pterostichus migliaccioi species	accepted (GBIF)	No associated record in PBDB
Pterostichus militaris species	accepted (GBIF)	No associated record in PBDB
Pterostichus minax species	accepted (GBIF)	No associated record in PBDB
Pterostichus ming species	accepted (GBIF)	No associated record in PBDB
Pterostichus minor species	accepted (GBIF) listed (PBDB)		4	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus minutulus ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Pterostichus mirificus species	accepted (GBIF)	No associated record in PBDB
Pterostichus miroshnikovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus miroslavi species	accepted (GBIF)	No associated record in PBDB
Pterostichus mirus species	accepted (GBIF)	No associated record in PBDB
Pterostichus miscellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mitoyamanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus miyamai species	accepted (GBIF)	No associated record in PBDB
Pterostichus mizunoyai species	accepted (GBIF)	No associated record in PBDB
Pterostichus modicellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus moestus species	accepted (GBIF)	No associated record in PBDB
Pterostichus molestus ✝ species	accepted (GBIF) listed (PBDB)		1	33.9 Ma Oligocene	33.9 Ma Oligocene
Pterostichus molopsoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus monostigma species	accepted (GBIF)	No associated record in PBDB
Pterostichus montanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus monticoloides species	accepted (GBIF)	No associated record in PBDB
Pterostichus montigena species	accepted (GBIF)	No associated record in PBDB
Pterostichus montisdeorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus morawitzianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus morio species	accepted (GBIF)	No associated record in PBDB
Pterostichus morionides species	accepted (GBIF)	No associated record in PBDB
Pterostichus mosaicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mucronatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus muehlfeldii species	accepted (GBIF)	No associated record in PBDB
Pterostichus muelleri species	accepted (GBIF)	No associated record in PBDB
Pterostichus muellermotzfeldi species	accepted (GBIF)	No associated record in PBDB
Pterostichus mujahedeeni species	accepted (GBIF)	No associated record in PBDB
Pterostichus mukdenensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus mulensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus multinodosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus multipunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus multiseta species	accepted (GBIF)	No associated record in PBDB
Pterostichus mundatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus mundus species	accepted (GBIF)	No associated record in PBDB
Pterostichus musashiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus mutoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus mutus species	accepted (GBIF)	No associated record in PBDB
Pterostichus nadari species	accepted (GBIF)	No associated record in PBDB
Pterostichus nakanei species	accepted (GBIF)	No associated record in PBDB
Pterostichus naokii species	accepted (GBIF)	No associated record in PBDB
Pterostichus napaea species	accepted (GBIF)	No associated record in PBDB
Pterostichus nasuensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus nasui species	accepted (GBIF)	No associated record in PBDB
Pterostichus necessarius species	accepted (GBIF)	No associated record in PBDB
Pterostichus neglectus species	accepted (GBIF)	No associated record in PBDB
Pterostichus negligens species	accepted (GBIF)	No associated record in PBDB
Pterostichus neilgaimani species	accepted (GBIF)	No associated record in PBDB
Pterostichus nemoralis species	accepted (GBIF)	No associated record in PBDB
Pterostichus neobrunneus species	accepted (GBIF)	No associated record in PBDB
Pterostichus nepalensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus nicaeensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus nigellatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus nigellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus niger species	accepted (GBIF) listed (PBDB)		2	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus nigripalpis species	accepted (GBIF)	No associated record in PBDB
Pterostichus nigrita species	accepted (GBIF) listed (PBDB)		2	2.59 Ma Pleistocene	0 Ma Extant
Pterostichus nigrocaeruleus species	accepted (GBIF)	No associated record in PBDB
Pterostichus nikodymi species	accepted (GBIF)	No associated record in PBDB
Pterostichus nimbatidius species	accepted (GBIF)	No associated record in PBDB
Pterostichus nishidai species	accepted (GBIF)	No associated record in PBDB
Pterostichus nishiyamai species	accepted (GBIF)	No associated record in PBDB
Pterostichus nivalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus nivicola species	accepted (GBIF)	No associated record in PBDB
Pterostichus noburai species	accepted (GBIF)	No associated record in PBDB
Pterostichus nodicornis species	accepted (GBIF)	No associated record in PBDB
Pterostichus noguchii species	accepted (GBIF)	No associated record in PBDB
Pterostichus nordqvisti species	accepted (GBIF)	No associated record in PBDB
Pterostichus novotnyorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus novus species	accepted (GBIF)	No associated record in PBDB
Pterostichus nowitzkii species	accepted (GBIF)	No associated record in PBDB
Pterostichus obesulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus oblongiusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus oblongoparallelus species	accepted (GBIF)	No associated record in PBDB
Pterostichus oblongopunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus obsidianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus occultus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ochoticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus odontocnemis species	accepted (GBIF)	No associated record in PBDB
Pterostichus odvarkai species	accepted (GBIF)	No associated record in PBDB
Pterostichus oenotrius species	accepted (GBIF)	No associated record in PBDB
Pterostichus ogaensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ohbayashii species	accepted (GBIF)	No associated record in PBDB
Pterostichus ohdaisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ohkawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus ohkurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus ohsawacavus species	accepted (GBIF)	No associated record in PBDB
Pterostichus okiensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus okutamae species	accepted (GBIF)	No associated record in PBDB
Pterostichus olafi species	accepted (GBIF)	No associated record in PBDB
Pterostichus olegi species	accepted (GBIF)	No associated record in PBDB
Pterostichus omogoensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ompoensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus opacipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ophryoderus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ordinarius species	accepted (GBIF)	No associated record in PBDB
Pterostichus ordinatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus orduensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus oregonus species	accepted (GBIF)	No associated record in PBDB
Pterostichus oreophilus species	accepted (GBIF)	No associated record in PBDB
Pterostichus orestes species	accepted (GBIF)	No associated record in PBDB
Pterostichus orientalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus orion species	accepted (GBIF)	No associated record in PBDB
Pterostichus orionis species	accepted (GBIF)	No associated record in PBDB
Pterostichus orman species	accepted (GBIF)	No associated record in PBDB
Pterostichus orobius species	accepted (GBIF)	No associated record in PBDB
Pterostichus ottomanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ovalipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ovaliphallus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ovoideus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pachinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus palekhai species	accepted (GBIF)	No associated record in PBDB
Pterostichus palgongsanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus palmi species	accepted (GBIF)	No associated record in PBDB
Pterostichus panticulatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pantomus species	accepted (GBIF)	No associated record in PBDB
Pterostichus panzeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus parasimilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus parkwon species	accepted (GBIF)	No associated record in PBDB
Pterostichus parnassius species	accepted (GBIF)	No associated record in PBDB
Pterostichus parviceps species	accepted (GBIF)	No associated record in PBDB
Pterostichus parvicollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus pasanauricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus patruelis species	accepted (GBIF)	No associated record in PBDB
Pterostichus paulini species	accepted (GBIF)	No associated record in PBDB
Pterostichus paulinoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus pavani species	accepted (GBIF)	No associated record in PBDB
Pterostichus pedemontanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus peilingi species	accepted (GBIF)	No associated record in PBDB
Pterostichus pemphredo species	accepted (GBIF)	No associated record in PBDB
Pterostichus peninsularis species	accepted (GBIF)	No associated record in PBDB
Pterostichus pensylvanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pentheri species	accepted (GBIF)	No associated record in PBDB
Pterostichus percontator species	accepted (GBIF)	No associated record in PBDB
Pterostichus pergracilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus perhoplites species	accepted (GBIF)	No associated record in PBDB
Pterostichus perisi species	accepted (GBIF)	No associated record in PBDB
Pterostichus perlamatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus perlutus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pertinax species	accepted (GBIF)	No associated record in PBDB
Pterostichus petulans species	accepted (GBIF)	No associated record in PBDB
Pterostichus pfizenmayeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus phaeopus species	accepted (GBIF)	No associated record in PBDB
Pterostichus phaeus species	accepted (GBIF)	No associated record in PBDB
Pterostichus phungaraziensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus piceolus species	accepted (GBIF)	No associated record in PBDB
Pterostichus piliferus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pilosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pinguedineus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pinguidineus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pithyusicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus planaticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus planctus species	accepted (GBIF)	No associated record in PBDB
Pterostichus planicola species	accepted (GBIF)	No associated record in PBDB
Pterostichus planiusculus species	accepted (GBIF)	No associated record in PBDB
Pterostichus planus species	accepted (GBIF)	No associated record in PBDB
Pterostichus platyderus species	accepted (GBIF)	No associated record in PBDB
Pterostichus platyops species	accepted (GBIF)	No associated record in PBDB
Pterostichus plesiomorphus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pohnerti species	accepted (GBIF)	No associated record in PBDB
Pterostichus polygenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ponticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus poppiusi species	accepted (GBIF)	No associated record in PBDB
Pterostichus poppiusianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus porcellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus potanini species	accepted (GBIF)	No associated record in PBDB
Pterostichus praedo species	accepted (GBIF)	No associated record in PBDB
Pterostichus pratti species	accepted (GBIF)	No associated record in PBDB
Pterostichus primus species	accepted (GBIF)	No associated record in PBDB
Pterostichus procax species	accepted (GBIF)	No associated record in PBDB
Pterostichus procephalus species	accepted (GBIF)	No associated record in PBDB
Pterostichus procerulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus properans species	accepted (GBIF)	No associated record in PBDB
Pterostichus protensiformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus protractus species	accepted (GBIF)	No associated record in PBDB
Pterostichus provincialis species	accepted (GBIF)	No associated record in PBDB
Pterostichus przewalskyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudoastutus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudobarbarae species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudodiversus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudoharmandi species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudojugivagus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudopachinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudopedius species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudopercus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudopilifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudoplatyderus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudoplatysma species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudorotundus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudosinensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus pseudostuxbergi species	accepted (GBIF)	No associated record in PBDB
Pterostichus pulchellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pulcher species	accepted (GBIF)	No associated record in PBDB
Pterostichus pumilio species	accepted (GBIF)	No associated record in PBDB
Pterostichus pumilus species	accepted (GBIF)	No associated record in PBDB
Pterostichus pumpellyi ✝ species	accepted (GBIF) listed (PBDB)		2	37.2 Ma Eocene	37.2 Ma Eocene
Pterostichus punctatissimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus punctibasalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus punctiger species	accepted (GBIF)	No associated record in PBDB
Pterostichus punctisternus species	accepted (GBIF)	No associated record in PBDB
Pterostichus punctiventris species	accepted (GBIF)	No associated record in PBDB
Pterostichus pusillus species	accepted (GBIF)	No associated record in PBDB
Pterostichus putus species	accepted (GBIF)	No associated record in PBDB
Pterostichus quadraticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus quadrifoveolatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus quadriimpressus species	accepted (GBIF)	No associated record in PBDB
Pterostichus quelpartensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus raptor species	accepted (GBIF)	No associated record in PBDB
Pterostichus raraensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rasilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus regeli species	accepted (GBIF)	No associated record in PBDB
Pterostichus reiseri species	accepted (GBIF)	No associated record in PBDB
Pterostichus reissi species	accepted (GBIF)	No associated record in PBDB
Pterostichus relictus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rengensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus restrictus species	accepted (GBIF)	No associated record in PBDB
Pterostichus reuteri species	accepted (GBIF)	No associated record in PBDB
Pterostichus rhaeticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rhanis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rhilensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rifensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus riparius species	accepted (GBIF)	No associated record in PBDB
Pterostichus ripensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus robustistylis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rolex species	accepted (GBIF)	No associated record in PBDB
Pterostichus rostratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rothi species	accepted (GBIF)	No associated record in PBDB
Pterostichus rotundangulus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rotundus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rousi species	accepted (GBIF)	No associated record in PBDB
Pterostichus rousianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rousselli species	accepted (GBIF)	No associated record in PBDB
Pterostichus rubripalpis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rudestriatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ruffoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus rufipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus rufitarsis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rufonitens species	accepted (GBIF)	No associated record in PBDB
Pterostichus rufopiceus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rugiceps species	accepted (GBIF)	No associated record in PBDB
Pterostichus rugosiceps species	accepted (GBIF)	No associated record in PBDB
Pterostichus rugosipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus rugosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rutenus species	accepted (GBIF)	No associated record in PBDB
Pterostichus rutilans species	accepted (GBIF)	No associated record in PBDB
Pterostichus ryomoensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sacheri species	accepted (GBIF)	No associated record in PBDB
Pterostichus sachtlebeni species	accepted (GBIF)	No associated record in PBDB
Pterostichus safonovae species	accepted (GBIF)	No associated record in PBDB
Pterostichus sagarmatha species	accepted (GBIF)	No associated record in PBDB
Pterostichus sakagamii species	accepted (GBIF)	No associated record in PBDB
Pterostichus sakuraii species	accepted (GBIF)	No associated record in PBDB
Pterostichus santostamangi species	accepted (GBIF)	No associated record in PBDB
Pterostichus sasajii species	accepted (GBIF)	No associated record in PBDB
Pterostichus satoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus satsumanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus satunini species	accepted (GBIF)	No associated record in PBDB
Pterostichus satyrus species	accepted (GBIF)	No associated record in PBDB
Pterostichus saueri species	accepted (GBIF)	No associated record in PBDB
Pterostichus saxicola species	accepted (GBIF)	No associated record in PBDB
Pterostichus sayanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus scalptus species	accepted (GBIF)	No associated record in PBDB
Pterostichus schaschli species	accepted (GBIF)	No associated record in PBDB
Pterostichus schmidtii species	accepted (GBIF)	No associated record in PBDB
Pterostichus schneideri species	accepted (GBIF)	No associated record in PBDB
Pterostichus schodaicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus schoenherri species	accepted (GBIF)	No associated record in PBDB
Pterostichus schoenmanni species	accepted (GBIF)	No associated record in PBDB
Pterostichus schrettenbrunneri species	accepted (GBIF)	No associated record in PBDB
Pterostichus schuelkei species	accepted (GBIF)	No associated record in PBDB
Pterostichus scitus species	accepted (GBIF)	No associated record in PBDB
Pterostichus scurra species	accepted (GBIF)	No associated record in PBDB
Pterostichus scutellaris species	accepted (GBIF)	No associated record in PBDB
Pterostichus scuticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sediorcus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sejunctus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sejungendus species	accepted (GBIF)	No associated record in PBDB
Pterostichus selmanni species	accepted (GBIF)	No associated record in PBDB
Pterostichus semirugosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus septemtrionalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus septentrionis species	accepted (GBIF)	No associated record in PBDB
Pterostichus seriatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus serripes species	accepted (GBIF)	No associated record in PBDB
Pterostichus seticeps species	accepted (GBIF)	No associated record in PBDB
Pterostichus setipes species	accepted (GBIF)	No associated record in PBDB
Pterostichus setosicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus setosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus seunglaki species	accepted (GBIF)	No associated record in PBDB
Pterostichus seungmoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus shaovuensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sharanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus shavrini species	accepted (GBIF)	No associated record in PBDB
Pterostichus shennongjianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus shibatai species	accepted (GBIF)	No associated record in PBDB
Pterostichus shiibanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus shilenkovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus shimizui species	accepted (GBIF)	No associated record in PBDB
Pterostichus shinbodakensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus shingarevi species	accepted (GBIF)	No associated record in PBDB
Pterostichus shirakamisan species	accepted (GBIF)	No associated record in PBDB
Pterostichus shirakamisanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus shirozui species	accepted (GBIF)	No associated record in PBDB
Pterostichus shojii species	accepted (GBIF)	No associated record in PBDB
Pterostichus shotaroi species	accepted (GBIF)	No associated record in PBDB
Pterostichus shulli species	accepted (GBIF)	No associated record in PBDB
Pterostichus sierranus species	accepted (GBIF)	No associated record in PBDB
Pterostichus similis species	accepted (GBIF)	No associated record in PBDB
Pterostichus simillimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sincerus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sinensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus singularis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sinicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sintanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus skoupyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus smetanai species	accepted (GBIF)	No associated record in PBDB
Pterostichus sodalicius species	accepted (GBIF)	No associated record in PBDB
Pterostichus sojaki species	accepted (GBIF)	No associated record in PBDB
Pterostichus solskyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus songoricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus soperi species	accepted (GBIF)	No associated record in PBDB
Pterostichus sotkaensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus spathifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus sphodriformis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sphodrinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus spiculifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus spinolae species	accepted (GBIF)	No associated record in PBDB
Pterostichus sponsor species	accepted (GBIF)	No associated record in PBDB
Pterostichus spornyi species	accepted (GBIF)	No associated record in PBDB
Pterostichus spraguei species	accepted (GBIF)	No associated record in PBDB
Pterostichus ssemenovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus stantonensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus stapedius species	accepted (GBIF)	No associated record in PBDB
Pterostichus starcki species	accepted (GBIF)	No associated record in PBDB
Pterostichus starckianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus staveni species	accepted (GBIF)	No associated record in PBDB
Pterostichus sterbai species	accepted (GBIF)	No associated record in PBDB
Pterostichus stictopleurus species	accepted (GBIF)	No associated record in PBDB
Pterostichus stoeckleini species	accepted (GBIF)	No associated record in PBDB
Pterostichus stomoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus straneellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus strasseri species	accepted (GBIF)	No associated record in PBDB
Pterostichus strenuus species	accepted (GBIF) listed (PBDB)		2	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus stricticollis species	accepted (GBIF)	No associated record in PBDB
Pterostichus strigipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus strigosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus stygicus species	accepted (GBIF) listed (PBDB)		2	2.59 Ma Pleistocene	0 Ma Extant
Pterostichus styx species	accepted (GBIF)	No associated record in PBDB
Pterostichus subabacipennis species	accepted (GBIF)	No associated record in PBDB
Pterostichus subacutus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subaeneus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subgibbus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subiasi species	accepted (GBIF)	No associated record in PBDB
Pterostichus subitus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sublaevis species	accepted (GBIF) listed (PBDB)		9	2.59 Ma Pleistocene	0 Ma Extant
Pterostichus subovatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subrugosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subsinuatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus subtilissimus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sudai species	accepted (GBIF)	No associated record in PBDB
Pterostichus sudrei species	accepted (GBIF)	No associated record in PBDB
Pterostichus suffusus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sugimurai species	accepted (GBIF)	No associated record in PBDB
Pterostichus sulcitarsis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sumondakensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus sungariensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus superciliosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus surgens species	accepted (GBIF)	No associated record in PBDB
Pterostichus suruganus species	accepted (GBIF)	No associated record in PBDB
Pterostichus sutschanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus svanicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus svetlanae species	accepted (GBIF)	No associated record in PBDB
Pterostichus swaneticus species	accepted (GBIF)	No associated record in PBDB
Pterostichus syleus species	accepted (GBIF)	No associated record in PBDB
Pterostichus symmetricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus szekessyianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus szetschuanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tachongi species	accepted (GBIF)	No associated record in PBDB
Pterostichus taebaegsanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tahirai species	accepted (GBIF)	No associated record in PBDB
Pterostichus takadateyamanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus takahashii species	accepted (GBIF)	No associated record in PBDB
Pterostichus takaosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus taksonyis species	accepted (GBIF)	No associated record in PBDB
Pterostichus talibani species	accepted (GBIF)	No associated record in PBDB
Pterostichus tamarae species	accepted (GBIF)	No associated record in PBDB
Pterostichus tamsii species	accepted (GBIF)	No associated record in PBDB
Pterostichus tanakai species	accepted (GBIF)	No associated record in PBDB
Pterostichus tanakaorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus tanjae species	accepted (GBIF)	No associated record in PBDB
Pterostichus tantillus species	accepted (GBIF)	No associated record in PBDB
Pterostichus taoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus taradakensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tareumiut species	accepted (GBIF)	No associated record in PBDB
Pterostichus tarsalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tateishiyamanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tatianae species	accepted (GBIF)	No associated record in PBDB
Pterostichus tatricus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tenenbaumianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tenuimarginatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tenuis species	accepted (GBIF)	No associated record in PBDB
Pterostichus teretis species	accepted (GBIF)	No associated record in PBDB
Pterostichus testaceus species	accepted (GBIF)	No associated record in PBDB
Pterostichus thanglaensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus theeli species	accepted (GBIF)	No associated record in PBDB
Pterostichus thorectes species	accepted (GBIF)	No associated record in PBDB
Pterostichus thorectoides species	accepted (GBIF)	No associated record in PBDB
Pterostichus thunbergi species	accepted (GBIF)	No associated record in PBDB
Pterostichus tichomirovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus tienmushanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tiliaceoradix species	accepted (GBIF)	No associated record in PBDB
Pterostichus todai species	accepted (GBIF)	No associated record in PBDB
Pterostichus tokejii species	accepted (GBIF)	No associated record in PBDB
Pterostichus tokmakovae species	accepted (GBIF)	No associated record in PBDB
Pterostichus tokui species	accepted (GBIF)	No associated record in PBDB
Pterostichus toledanoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus tomensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tonkinensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tottoriensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus toyodai species	accepted (GBIF)	No associated record in PBDB
Pterostichus transversalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus triangularis species	accepted (GBIF)	No associated record in PBDB
Pterostichus trinarius species	accepted (GBIF)	No associated record in PBDB
Pterostichus trinitensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus triseriatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tristis species	accepted (GBIF)	No associated record in PBDB
Pterostichus troglophilus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tropicalis species	accepted (GBIF)	No associated record in PBDB
Pterostichus trubilini species	accepted (GBIF)	No associated record in PBDB
Pterostichus truncatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tschitscherinianus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tsukubasanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tsurugaensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tsurugiyamanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tuberculiger species	accepted (GBIF)	No associated record in PBDB
Pterostichus tuberculofemoratus species	accepted (GBIF)	No associated record in PBDB
Pterostichus tuberifer species	accepted (GBIF)	No associated record in PBDB
Pterostichus tukchensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus tulobalu species	accepted (GBIF)	No associated record in PBDB
Pterostichus tumescens species	accepted (GBIF)	No associated record in PBDB
Pterostichus tundrae species	accepted (GBIF) listed (PBDB)		6	2.59 Ma Pleistocene	0 Ma Extant
Pterostichus turanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus turcomannicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus uchiyamai species	accepted (GBIF)	No associated record in PBDB
Pterostichus uedaorum species	accepted (GBIF)	No associated record in PBDB
Pterostichus uenoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus ulubeyensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus unctulatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus uralensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus urengaicus species	accepted (GBIF)	No associated record in PBDB
Pterostichus ussuriensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus vagepunctatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus validiceps species	accepted (GBIF)	No associated record in PBDB
Pterostichus validior species	accepted (GBIF)	No associated record in PBDB
Pterostichus validus species	accepted (GBIF)	No associated record in PBDB
Pterostichus vandykei species	accepted (GBIF)	No associated record in PBDB
Pterostichus variabilis species	accepted (GBIF)	No associated record in PBDB
Pterostichus variicornis species	accepted (GBIF)	No associated record in PBDB
Pterostichus variolatus species	accepted (GBIF)	No associated record in PBDB
Pterostichus variseta species	accepted (GBIF)	No associated record in PBDB
Pterostichus vecors species	accepted (GBIF)	No associated record in PBDB
Pterostichus ventralis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ventricosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus vermiculosus species	accepted (GBIF)	No associated record in PBDB
Pterostichus vernalis species	accepted (GBIF) listed (PBDB)		5	0.78 Ma Pleistocene	0 Ma Extant
Pterostichus vetustus ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Pterostichus vicinus species	accepted (GBIF)	No associated record in PBDB
Pterostichus violaceotinctus species	accepted (GBIF)	No associated record in PBDB
Pterostichus virescens species	accepted (GBIF)	No associated record in PBDB
Pterostichus viridellus species	accepted (GBIF)	No associated record in PBDB
Pterostichus vladivostokensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus vseteckai species	accepted (GBIF)	No associated record in PBDB
Pterostichus walcotti ✝ species	accepted (GBIF) listed (PBDB)		3	37.2 Ma Eocene	37.2 Ma Eocene
Pterostichus walteri species	accepted (GBIF)	No associated record in PBDB
Pterostichus watanabei species	accepted (GBIF)	No associated record in PBDB
Pterostichus weigeli species	accepted (GBIF)	No associated record in PBDB
Pterostichus weiperti species	accepted (GBIF)	No associated record in PBDB
Pterostichus wenxianensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus wittmeri species	accepted (GBIF)	No associated record in PBDB
Pterostichus woodi species	accepted (GBIF)	No associated record in PBDB
Pterostichus woongbii species	accepted (GBIF)	No associated record in PBDB
Pterostichus woronowi species	accepted (GBIF)	No associated record in PBDB
Pterostichus wrasei species	accepted (GBIF)	No associated record in PBDB
Pterostichus xatartii species	accepted (GBIF)	No associated record in PBDB
Pterostichus xilingensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus yahikosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus yakushimanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus yamajii species	accepted (GBIF)	No associated record in PBDB
Pterostichus yamauchii species	accepted (GBIF)	No associated record in PBDB
Pterostichus yamizosanus species	accepted (GBIF)	No associated record in PBDB
Pterostichus yanoi species	accepted (GBIF)	No associated record in PBDB
Pterostichus yatsuensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ybousqueti species	accepted (GBIF)	No associated record in PBDB
Pterostichus yokohamae species	accepted (GBIF)	No associated record in PBDB
Pterostichus yorikoae species	accepted (GBIF)	No associated record in PBDB
Pterostichus yoritomus species	accepted (GBIF)	No associated record in PBDB
Pterostichus yosemitensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus yoshidai species	accepted (GBIF)	No associated record in PBDB
Pterostichus yoshikawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus yoshizawai species	accepted (GBIF)	No associated record in PBDB
Pterostichus yulongshanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus yunnanensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus yvanii species	accepted (GBIF)	No associated record in PBDB
Pterostichus zamotajlovi species	accepted (GBIF)	No associated record in PBDB
Pterostichus zhejiangensis species	accepted (GBIF)	No associated record in PBDB
Pterostichus ziegleri species	accepted (GBIF)	No associated record in PBDB
Pterostichus zoiai species	accepted (GBIF)	No associated record in PBDB
Pterostichus zolotarewi species	accepted (GBIF)	No associated record in PBDB
Pterostichus zyzzovi species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Peryphes Casey, 1920	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Peryphes*
Parahaptoderus Jeanne, 1969	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Parahaptoderus*
Sinosteropus Sciaky, 1994	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sinosteropus*
Paleocorax Ortuno, 1990	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paleocorax*
Jeannelinus Schatzmayr, 1943	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Jeannelinus*
Orsonjohnsonus Hatch, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Orsonjohnsonus*
Cophosus Dejean, 1821	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cophosus*
Aphaon Heyden, 1906	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aphaon*
Bothriopterus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Bothriopterus*
Morphnosoma Lucnik, 1915	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Morphnosoma*
Phaenorrhaphis Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Phaenorrhaphis*
Paralianoe Ishida, 1958	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paralianoe*
Adelosia Stephens, 1835	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Adelosia*
Biargutor Novoa, 1979	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Biargutor*
Corax Putzeys, 1846	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Corax*
Morphaptoderus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Morphaptoderus*
Feronidius Jeannel, 1942	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Feronidius*
Coptosus Gistl, 1830	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Coptosus*
Aello Gozis, 1882	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aello*
Iberophilus Jeanne, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Iberophilus*
Eurypessus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eurypessus*
Eolargarus Tschitscherine, 1896-1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eolargarus*
Abaxodes Gistel, 1857	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Abaxodes*
Platysma Bonelli, 1810	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platysma*
Omaseidus Jeannel, 1942	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omaseidus*
Rhagodus Motschoulsky, 1866	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhagodus*
Tritrichis Andrewes, 1937	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Tritrichis*
Euhaptorderus Jeanne, 1969	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Euhaptorderus*
Wraseiellus Shi & Sciaky, 2013	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Wraseiellus*
Feronina Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Feronina*
Pseudorites Ganglbauer, 1891	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudorites*
Steropinus Lucnik, 1916	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Steropinus*
Ragadus Motschoulsky, 1865	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ragadus*
Sterocorax Ortuno, 1990	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sterocorax*
Leptoferonia Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Leptoferonia*
Holciophorus Leconte, 1853	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Holciophorus*
Haptoderus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haptoderus*
Nehaptoderus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Nehaptoderus*
Dysidius Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Dysidius*
Myosodus Megerle & Steven, 1824	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Myosodus*
Ethira Andrewes, 1936	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ethira*
Koreonialoe Park & Kwon, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Koreonialoe*
Bryobius de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Bryobius*
Pseudomasoeus Desmarest, 1851-22	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudomasoeus*
Paralagarus Lucnik, 1922	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paralagarus*
Amolopsa Strand, 1936	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amolopsa*
Bryabius de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Bryabius*
Amolops Tschitscherin, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amolops*
Melanius Bonelli, 1810	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Melanius*
Omasoeus Desmarest, 1851-22	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omasoeus*
Unitrichus Sciaky, 1997	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Unitrichus*
Omazeus Laporte, 1834	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omazeus*
Brachystilus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Brachystilus*
Jedlickaia Sciaky, 1997	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Jedlickaia*
Melvilleus Ball, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Melvilleus*
Callipterus Agassiz, 1846	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Callipterus*
Glyptopterus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Glyptopterus*
Phonias Des Gozis, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Phonias*
Hammatomerus de Chaudoir, 1868	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hammatomerus*
Pseudosteropus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudosteropus*
Platypterus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platypterus*
Fukienostichus Lafer, 1983	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Fukienostichus*
Steropus Stephens, 1828	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Steropus*
Anomostichus Sciaky, 1995	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Anomostichus*
Haplocoelus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haplocoelus*
Pyreneorites Jeannel, 1937	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pyreneorites*
Circinatus Sciaky, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Circinatus*
Devilleinus Schatzmayr, 1943	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Devilleinus*
Haplomaseus Reitter, 1896	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haplomaseus*
Pterostichus Stephens, 1827	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pterostichus*
Steroperis Shilenkov, 1995	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Steroperis*
Brachystylus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Brachystylus*
Sinoreophilus Sciaky, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sinoreophilus*
Nialoe Tanaka, 1958	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Nialoe*
Japeris Sciaky, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Japeris*
Phaenoraphis Tschitscherin, 1901	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Phaenoraphis*
Lyperopherus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lyperopherus*
Hypherpes Eschscholtz, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hypherpes*
Paraferonia Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Paraferonia*
Cosciniopterus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cosciniopterus*
Phaon Heyden, 1883	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Phaon*
Daisenialoe Nakane & Straneo, 1979	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Daisenialoe*
Lyrothorax de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lyrothorax*
Pseudoferonina Ball, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudoferonina*
Oreophilus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Oreophilus*
Agonodemus de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonodemus*
Agonoderus Gemminger & Harold, 1868	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonoderus*
Micromaseus Desbrochers, 1906	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Micromaseus*
Morphohaptoderus Tschitscherin, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Morphohaptoderus*
Omasaeus Perty, 1833	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omasaeus*
Steroderus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Steroderus*
Parasteropus Reitter, 1908	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Parasteropus*
Americomaseus Csiki, 1930	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Americomaseus*
Aachnoideus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aachnoideus*
Jeannelinus Jeanne, 1965	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Jeannelinus*
Haploderus Torre, 1877	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haploderus*
Gastrostricta Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Gastrostricta*
Cheropus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cheropus*
Hysherpes Desmarest, 1851-22	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hysherpes*
Rhagadus Motschoulsky, 1865	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhagadus*
Micromaseus Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Micromaseus*
Cylindrocharis Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cylindrocharis*
Gonoderus Motschoulsky, 1859	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Gonoderus*
Badistrinus Motschoulsky, 1865	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Badistrinus*
Omaseulus Lucnik, 1929	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omaseulus*
Petrophilus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Petrophilus*
Cophus Agassiz, 1846	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cophus*
Biphonias Jeanne, 1988	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Biphonias*
Pseudolagarus Jedlicka, 1956	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudolagarus*
Haptoderotapinus Jedlicka, 1930	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haptoderotapinus*
Agonodemus Reitter, 1896	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Agonodemus*
Lyperosomus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lyperosomus*
Omascus Harris, 1828	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omascus*
Pledarus Motschoulsky, 1865	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pledarus*
Galapterus Berlov & Plutenko, 1997	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Galapterus*
Glyptodactylus Cottes, 1870	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Glyptodactylus*
Natalianoe Berlov & Plutenko, 1997	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Natalianoe*
Georgebaalus Habu, 1985	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Georgebaalus*
Dasalus Motschulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Dasalus*
Iberoderus Jeanne, 1969	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Iberoderus*
Logarus Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Logarus*
Carllindrothius Habu, 1985	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Carllindrothius*
Monoferonia Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Monoferonia*
Pseudadelosia Tschitscherin, 1889	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudadelosia*
Euferonia Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Euferonia*
Haptotapinus Reitter, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haptotapinus*
Nemotopterus Berlov, 2000	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Nemotopterus*
Oreolyperus Tschitscherin, 1901	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Oreolyperus*
Cheporus Dahl, 1823	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cheporus*
Pseudoomaseus Motschoulsky, 1869	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudoomaseus*
Plectes Fischer de Waldheim, 1821	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Plectes*
Metamelanius Tschitscherin, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Metamelanius*
Plectus Sturm, 1843	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Plectus*
Peudhaptoderus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Peudhaptoderus*
Metallophilus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Metallophilus*
Lagarus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lagarus*
Lyperus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lyperus*
Calopterus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Calopterus*
Pseudorthomus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudorthomus*
Asioplatysma Kryzhanovskii, 1968	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Asioplatysma*
Melanius Dejean, 1828	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Melanius*
Pseudolagarus Lucnik, 1922	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudolagarus*
Sphodroferonia Kasahara & Ito, 1987	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sphodroferonia*
Neohaptoderus Tschitscherin, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Neohaptoderus*
Ploctes Sturm, 1843	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ploctes*
Platysma LeConte, 1873	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platysma*
Pseudargutor Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudargutor*
Orthomus Motschulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Orthomus*
Refonia Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Refonia*
Orites Schaum, 1858	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Orites*
Tschitscherinea Berg, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Tschitscherinea*
Adelopterus Reitter, 1886	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Adelopterus*
Platisma Hope, 1831	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platisma*
Parargutor Casey, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Parargutor*
Anilloferonia Van Dyke, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Anilloferonia*
Moritapterus Berlov, 2000	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Moritapterus*
Eurythorax Tschitscherin, 1889	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eurythorax*
Platysma Dejean, 1821	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platysma*
Euryperis Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Euryperis*
Epinialoe Nakane, 1979	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Epinialoe*
Pseudomaseus Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudomaseus*
Psychobius de Chaudoir, 1838	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Psychobius*
Alecto Heyden, 1883	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Alecto*
Oreoplatysma Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Oreoplatysma*
Falsargutor Kryzhanovskij, 1983	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Falsargutor*
Feroperis Lafer, 1979	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Feroperis*
Hapterotapinus Csiki, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hapterotapinus*
Micronialoe Park, Kwon & Lafer, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Micronialoe*
Eosteropus Tschitscherin, 1902	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eosteropus*
Platypterinus Reitter, 1908	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platypterinus*
Eunema Tschitscherin, 1897	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eunema*
Eurymelanius Reitter, 1896	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eurymelanius*
Iberopus Ortuno, 1990	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Iberopus*
Cheporus Latreille, 1829	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cheporus*
Steropidius Jeanne, 1989	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Steropidius*
Platyderus Kirby, 1837	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platyderus*
Luperopherus Motschoulsky, 1869	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Luperopherus*
Gutta Wrase & Schmidt, 2006	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Gutta*
Pseudethira Sciaky, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudethira*
Aulacotarsus Reiche, 1872	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Aulacotarsus*
Megabea Sciaky, 1997	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Megabea*
Platysma Samouelle, 1819	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platysma*
Abea Morita, 1992	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Abea*
Trichomolops Andrewes, 1939	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Trichomolops*
Pseudohaptoderus Tschitscherin, 1888	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudohaptoderus*
Arguta Lucas, 1846	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Arguta*
Eurythoracana Strand, 1936	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Eurythoracana*
Pseudocryobius Motschoulsky, 1850	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudocryobius*
Lianoe Gozis, 1882	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lianoe*
Omaseus Dejean, 1821	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Omaseus*
Rhagadulus Tschitscherin, 1897	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rhagadulus*

References

Bousquet, Yves, and André Larochelle, 1993: Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, no. 167. 397. - via Integrated Taxonomic Information System (ITIS)

Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722. - via Integrated Taxonomic Information System (ITIS)

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

WoRMS (Mar 2013) - via The Interim Register of Marine and Nonmarine Genera

as per family (nonmarine except where stated) - via The Interim Register of Marine and Nonmarine Genera

Fauna Europaea (May 2010) - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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