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Blethisa

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Description

Blethisa is a genus of ground beetle native to the Palearctic. It contains the following species:

From Wikipedia article at https://en.wikipedia.org/wiki/Blethisa, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	4988206	https://www.gbif.org/species/4988206
PaleoBioDB ID (PBDB)	193884	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=193884

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Blethisa

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
subfamily	Elaphrinae	Latreille 1802
genus	Blethisa	Bonelli 1810

Scientific Name

Blethisa Bonelli, 1810

Name Published In

Bonelli, Franco A. 1810. Observations entomologiques. Première partie (cicindélètes et portion des carabiques). Félix Galletti, Turin.: 1-58; Tabula synoptica.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Blethisa	genus	belongs to Carabiques	stated with evidence	Dejean, 1821
Blethisa	genus	belongs to Elaphrini	stated with evidence	LeConte, 1861
Blethisa	genus	belongs to Carabidae	stated without evidence	Kiselev and Nazarov, 2009
Blethisa	genus	belongs to Elaphrini	stated with evidence	Tamutis et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

4 (80%)

First Recorded Appearance

11.6 - 5.3 Ma
Miocene

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

F. A. Bonelli. 1810. Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques)

Wikipedia

https://en.wikipedia.org/wiki/Blethisa

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Blethisa catenaria species	accepted (GBIF) listed (PBDB)		5	2.59 Ma Pleistocene	0 Ma Extant
Blethisa eschscholtzii species	accepted (GBIF)	No associated record in PBDB
Blethisa hudsonica species	accepted (GBIF)	No associated record in PBDB
Blethisa inexpectata species	accepted (GBIF) listed (PBDB)		1	11.6 Ma Miocene	0 Ma Extant
Blethisa julii species	accepted (GBIF)	No associated record in PBDB
Blethisa multipunctata species	accepted (GBIF) listed (PBDB)		3	2.59 Ma Pleistocene	0 Ma Extant
Blethisa oregonenis species	accepted (GBIF)	No associated record in PBDB
Blethisa oregonensis species	accepted (GBIF)	No associated record in PBDB
Blethisa pleistocenica ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Blethisa quadricollis species	accepted (GBIF)	No associated record in PBDB
Blethisa tuberculata species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Blethisa Dejean, 1826	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Blethisa*
Raphiona Fischer von Waldheim, 1829	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Raphiona*
Helobium Leach, 1815	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Helobium*

Synonymy List

Year	Name and Author
1810	Blethisa Bonelli
1821	Blethisa Dejean p. 18
1861	Blethisa LeConte p. 7
1983	Blethisa Goulet
2009	Blethisa Kiselev and Nazarov
2011	Blethisa Tamutis et al.

References

Bonelli F. A. , Observations Entomologiques. Première Partie (Cicindélètes et Portion des Carabiques), 1-58

Dejean P. F. M. , Catalogue de la collection de Coléoptères de M. le Baron Dejean, 1-136

Goulet H. (1983) The genera of Holarctic Elaphrini and species of Elaphrus Fabricius (Coleoptera: Carabidae): Classification, phylogeny and zoogeography, Quaestiones Entomologicae 19, 219-482

Kiselev S. V., Nazarov V. I. (2009) Late Cenozoic insects of northern Eurasia, Paleontological Journal 43 7, 732-850

Tamutis V., Tamuté B., et al (2011) A catalogue of Lithuanian beetles (Insecta, Coleoptera), ZooKeys 121, 1-494

Goulet H, Smetana A (1983) A new species of Blethisa Bonelli from Alaska, with proposed phylogeny, biogeography and key to known species (Coleoptera: Carabidae). The Canadian Entomologist 115: 551-558. - via Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition.

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

as per family (nonmarine except where stated) - via The Interim Register of Marine and Nonmarine Genera

Erwin, Terry L., 2011: null. A Treatise on the Western Hemisphere Caraboidea (Coleoptera): Their classification, distributions, and ways of life, volume III. Carabidae - Loxomeriformes, Melaeniformes. 412. - via Integrated Taxonomic Information System (ITIS)

Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722. - via Integrated Taxonomic Information System (ITIS)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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