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Leptoconops

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Description

Leptoconops (black gnat) is a midge genus in the family Ceratopogonidae. It has a mostly tropical or subtropical distribution worldwide, but some species occur as far north as Moscow region in Russia and the Yukon Territory in Canada.

From Wikipedia article at https://en.wikipedia.org/wiki/Leptoconops, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	1633120	https://www.gbif.org/species/1633120
PaleoBioDB ID (PBDB)	204205	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=204205

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : Leptoconops

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Mecopterida
superorder	Antliophora
order	Diptera	Linnaeus 1758
infraorder	Culicomorpha
superfamily	Chironomoidea	Macquart 1838
family	Ceratopogonidae	Newman 1834
subfamily	Leptoconopinae	Noe 1907
genus	Leptoconops	Skuse 1889

Scientific Name

Leptoconops Skuse, 1889

Name Published In

Proc. Linn. Soc. N. S. Wales, (2) 4

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Leptoconops	genus	belongs to Ceratopogoninae	stated with evidence	Kieffer, 1906
Leptoconops	genus	belongs to Leptoconopinae	stated with evidence	Szadziewski, 1988
Leptoconops	genus	belongs to Ceratopogonidae	stated without evidence	Borkent, 1997
Leptoconops	genus	belongs to Leptoconopinae	stated with evidence	Borkent and Wirth, 1997

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

2 (9.1%)

First Recorded Appearance

129 - 125 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

A. Borkent. 1997. Upper and Lower Cretaceous Biting Midges (Ceratopogonidae: Diptera) from Hungarian and Austrian Amber and the Koonwarra Fossil Bed of Australia. Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 249:1-10

Wikipedia

https://en.wikipedia.org/wiki/Leptoconops

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Leptoconops (Leptoconops) subgenus	listed (PBDB)		27	105 Ma Early/Lower Cretaceous	0 Ma Extant
Leptoconops (Palaeoconops) ✝ subgenus	listed (PBDB)		2	129 Ma Early/Lower Cretaceous	129 Ma Early/Lower Cretaceous
Leptoconops acer species	accepted (GBIF)	No associated record in PBDB
Leptoconops albiventris species	accepted (GBIF)	No associated record in PBDB
Leptoconops algeriensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops americanus species	accepted (GBIF)	No associated record in PBDB
Leptoconops amplifemoralis species	accepted (GBIF)	No associated record in PBDB
Leptoconops amplificatus species	accepted (GBIF)	No associated record in PBDB
Leptoconops andersoni species	accepted (GBIF)	No associated record in PBDB
Leptoconops antiquus species	accepted (GBIF)	No associated record in PBDB
Leptoconops arnaudi species	accepted (GBIF)	No associated record in PBDB
Leptoconops asilomar species	accepted (GBIF)	No associated record in PBDB
Leptoconops atchleyi species	accepted (GBIF)	No associated record in PBDB
Leptoconops auster species	accepted (GBIF)	No associated record in PBDB
Leptoconops australiensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops aviarum species	accepted (GBIF)	No associated record in PBDB
Leptoconops bahreinensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops bassoi species	accepted (GBIF)	No associated record in PBDB
Leptoconops belkini species	accepted (GBIF)	No associated record in PBDB
Leptoconops bequaerti species	accepted (GBIF)	No associated record in PBDB
Leptoconops bezzii species	accepted (GBIF)	No associated record in PBDB
Leptoconops bidentatus species	accepted (GBIF)	No associated record in PBDB
Leptoconops binangulus species	accepted (GBIF)	No associated record in PBDB
Leptoconops borealis species	accepted (GBIF)	No associated record in PBDB
Leptoconops boreus species	accepted (GBIF)	No associated record in PBDB
Leptoconops brasiliensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops brevistylus species	accepted (GBIF)	No associated record in PBDB
Leptoconops bullsbrookensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops bundyensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops burmiticus species	accepted (GBIF)	No associated record in PBDB
Leptoconops californiensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops camelorum species	accepted (GBIF)	No associated record in PBDB
Leptoconops capensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops carteri species	accepted (GBIF)	No associated record in PBDB
Leptoconops casali species	accepted (GBIF)	No associated record in PBDB
Leptoconops catawbae species	accepted (GBIF)	No associated record in PBDB
Leptoconops caucasicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops chilensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops chinensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops clava species	accepted (GBIF)	No associated record in PBDB
Leptoconops conulus species	accepted (GBIF)	No associated record in PBDB
Leptoconops copiosus species	accepted (GBIF)	No associated record in PBDB
Leptoconops curvachelus species	accepted (GBIF)	No associated record in PBDB
Leptoconops daugeroni species	accepted (GBIF)	No associated record in PBDB
Leptoconops demeilloni species	accepted (GBIF)	No associated record in PBDB
Leptoconops dissimilis species	accepted (GBIF)	No associated record in PBDB
Leptoconops dixi species	accepted (GBIF)	No associated record in PBDB
Leptoconops doyeni species	accepted (GBIF)	No associated record in PBDB
Leptoconops ellenbergeri species	accepted (GBIF)	No associated record in PBDB
Leptoconops endialis species	accepted (GBIF)	No associated record in PBDB
Leptoconops exspectator species	accepted (GBIF)	No associated record in PBDB
Leptoconops flaviventris species	accepted (GBIF)	No associated record in PBDB
Leptoconops floridensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops foleyi species	accepted (GBIF)	No associated record in PBDB
Leptoconops fortipalpus species	accepted (GBIF)	No associated record in PBDB
Leptoconops foulki species	accepted (GBIF)	No associated record in PBDB
Leptoconops freeborni species	accepted (GBIF)	No associated record in PBDB
Leptoconops fuscipennis species	accepted (GBIF)	No associated record in PBDB
Leptoconops gallicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops golanensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops grandis species	accepted (GBIF)	No associated record in PBDB
Leptoconops gravesi species	accepted (GBIF)	No associated record in PBDB
Leptoconops halophilus species	accepted (GBIF)	No associated record in PBDB
Leptoconops hamariensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops harrisoni species	accepted (GBIF)	No associated record in PBDB
Leptoconops hutsoni species	accepted (GBIF)	No associated record in PBDB
Leptoconops hyalinipennis species	accepted (GBIF)	No associated record in PBDB
Leptoconops indicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops interruptus species	accepted (GBIF)	No associated record in PBDB
Leptoconops irritans species	accepted (GBIF)	No associated record in PBDB
Leptoconops kerteszi species	accepted (GBIF)	No associated record in PBDB
Leptoconops knowltoni species	accepted (GBIF)	No associated record in PBDB
Leptoconops lacteipennis species	accepted (GBIF)	No associated record in PBDB
Leptoconops latibulorum species	accepted (GBIF)	No associated record in PBDB
Leptoconops laurae species	accepted (GBIF)	No associated record in PBDB
Leptoconops leptorhynchus species	accepted (GBIF)	No associated record in PBDB
Leptoconops linleyi species	accepted (GBIF)	No associated record in PBDB
Leptoconops lisbonnei species	accepted (GBIF)	No associated record in PBDB
Leptoconops longicornis species	accepted (GBIF)	No associated record in PBDB
Leptoconops lucidus species	accepted (GBIF)	No associated record in PBDB
Leptoconops macfiei species	accepted (GBIF)	No associated record in PBDB
Leptoconops mackerrasae species	accepted (GBIF)	No associated record in PBDB
Leptoconops melanderi species	accepted (GBIF)	No associated record in PBDB
Leptoconops mellori species	accepted (GBIF)	No associated record in PBDB
Leptoconops minutus species	accepted (GBIF)	No associated record in PBDB
Leptoconops mohavensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops montanus species	accepted (GBIF)	No associated record in PBDB
Leptoconops montigena species	accepted (GBIF)	No associated record in PBDB
Leptoconops mooloolabaensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops myanmaricus species	accepted (GBIF)	No associated record in PBDB
Leptoconops myersi species	accepted (GBIF)	No associated record in PBDB
Leptoconops nachitschevanicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops nevilli species	accepted (GBIF)	No associated record in PBDB
Leptoconops nicolayi species	accepted (GBIF)	No associated record in PBDB
Leptoconops nigripes species	accepted (GBIF)	No associated record in PBDB
Leptoconops nipponensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops nivalis species	accepted (GBIF)	No associated record in PBDB
Leptoconops noei species	accepted (GBIF)	No associated record in PBDB
Leptoconops nosopheris species	accepted (GBIF)	No associated record in PBDB
Leptoconops obscurus species	accepted (GBIF)	No associated record in PBDB
Leptoconops panamensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops parvichela species	accepted (GBIF)	No associated record in PBDB
Leptoconops patagoniensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops pavlovskyi species	accepted (GBIF)	No associated record in PBDB
Leptoconops peneti species	accepted (GBIF)	No associated record in PBDB
Leptoconops petrocchiae species	accepted (GBIF)	No associated record in PBDB
Leptoconops primaevus species	accepted (GBIF)	No associated record in PBDB
Leptoconops pseudosetosifrons species	accepted (GBIF)	No associated record in PBDB
Leptoconops pugnax species	accepted (GBIF)	No associated record in PBDB
Leptoconops qinghaiensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops reesi species	accepted (GBIF)	No associated record in PBDB
Leptoconops rhodesiensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops ricardoi species	accepted (GBIF)	No associated record in PBDB
Leptoconops riverinaensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops rossi species	accepted (GBIF)	No associated record in PBDB
Leptoconops rovnensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops setosifrons species	accepted (GBIF)	No associated record in PBDB
Leptoconops shangweni species	accepted (GBIF)	No associated record in PBDB
Leptoconops siamensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops sibiricus species	accepted (GBIF)	No associated record in PBDB
Leptoconops smeei species	accepted (GBIF)	No associated record in PBDB
Leptoconops spinosifrons species	accepted (GBIF)	No associated record in PBDB
Leptoconops stygius species	accepted (GBIF)	No associated record in PBDB
Leptoconops sublettei species	accepted (GBIF)	No associated record in PBDB
Leptoconops subrossicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops succineus species	accepted (GBIF)	No associated record in PBDB
Leptoconops tarimensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops tenebrostigmatus species	accepted (GBIF)	No associated record in PBDB
Leptoconops tibetensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops torrens species	accepted (GBIF)	No associated record in PBDB
Leptoconops tuotuohea species	accepted (GBIF)	No associated record in PBDB
Leptoconops turkmenicus species	accepted (GBIF)	No associated record in PBDB
Leptoconops umbellifer species	accepted (GBIF)	No associated record in PBDB
Leptoconops vargasi species	accepted (GBIF)	No associated record in PBDB
Leptoconops venezuelensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops vexans species	accepted (GBIF)	No associated record in PBDB
Leptoconops werneri species	accepted (GBIF)	No associated record in PBDB
Leptoconops whitseli species	accepted (GBIF)	No associated record in PBDB
Leptoconops woodhilli species	accepted (GBIF)	No associated record in PBDB
Leptoconops xuthosceles species	accepted (GBIF)	No associated record in PBDB
Leptoconops yixini species	accepted (GBIF)	No associated record in PBDB
Leptoconops yunhsienensis species	accepted (GBIF)	No associated record in PBDB
Leptoconops zherikhini species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Protersesthes Kieffer, 1921	GBIF	genus	Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : *Protersesthes*
Schizoconops Kieffer, 1918	GBIF	genus	Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : *Schizoconops*
Centrotypus Grassi, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : *Centrotypus*
Mycterotypus Noe, 1905	GBIF	genus	Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : *Mycterotypus*
Tersesthes Townsend, 1893	GBIF	genus	Animalia : Arthropoda : Insecta : Diptera : Ceratopogonidae : *Tersesthes*

Synonymy List

Year	Name and Author
1889	Leptoconops Skuse
1906	Leptoconops Kieffer p. 48
1988	Leptoconops Szadziewski p. 232
1990	Leptoconops Szadziewski p. 79
1996	Leptoconops Szadziewski p. 48
1997	Leptoconops Borkent p. 4
1997	Leptoconops Borkent and Wirth p. 16

Year	Name and Author
2011	Leptoconops Sontag and Szadziewski p. 780
2013	Leptoconops Borkent p. 12
2014	Leptoconops Choufani et al. p. 35
2015	Leptoconops Szadziewski et al. p. 256
2015	Leptoconops Szadziewski et al. p. 557
2018	Leptoconops Pielowska et al. p. 241

References

Borkent A. (1997) Upper and Lower Cretaceous Biting Midges (Ceratopogonidae: Diptera) from Hungarian and Austrian Amber and the Koonwarra Fossil Bed of Australia, Stuttgarter Beiträge zur Naturkunde Serie B (Geologie und Paläontologie) 249, 1-10

Kieffer J. J. (1906) Diptera, Fam. Chironomidae, Genera Insectorum 42, 1-78

Szadziewski R. (1988) Biting midges (Diptera, Ceratopogonidae) from Baltic amber, Polskie Pismo Entomologiczne 57, 3-283

Szadziewski R. (1990) Biting midges (Insecta: Diptera: Ceratopogonidae) from Sakhalin amber, Prace Muzeum Ziemi 41, 77-82

Szadziewski R. (1996) Biting midges from Lower Cretaceous amber of Lebanon and Upper Cretaceous Siberian amber of Taimyr (Dipera, Ceratopogonidae), Studia Dipterologica 3, 23-86

Sontag E., Szadziewski R. (2011) Biting midges (Diptera: Ceratopogonidae) in Eocene Baltic amber from the Rovno region (Ukraine), Polish Journal of Entomology 80, 779-800

Borkent A. (2013) , World species of biting midges (Diptera: Ceratopogonidae)

Choufani J., Perrichot V., et al (2014) New biting midges (Diptera: Ceratopogonidae) in Late Cretaceous Vendean amber, Paleontological Contributions 10, 34-40

Szadziewski R., Gika W., et al (2015) A blood sucking biting midge from Upper Cretaceous Burmese amber with a key to the determination of fossil species in the relictual genus Leptoconops Skuse (Diptera: Ceratopogonidae), Cretaceous Research 54, 255-259

Szadziewski R., Ross A. J., et al (2015) Further records of biting midges (Diptera: Ceratopogonidae) from Upper Cretaceous Burmese amber (Myanmar), Cretaceous Research 52, 556-561

Pielowska A., Sontag E., et al (2018) Haematophagous arthropods in Baltic amber, Annales Zoologici 68, 237-249

Cheng, L. (Ed.). (1976). Marine insects. North-Holland Publishing Company: Amsterdam, The Netherlands. ISBN 0-444-11213-8. XII, 581 pp. - via World Register of Marine Species

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

SN2000 unverified - via The Interim Register of Marine and Nonmarine Genera

WoRMS (Mar 2013) - via The Interim Register of Marine and Nonmarine Genera

as per family - via The Interim Register of Marine and Nonmarine Genera

Proc. Linn. Soc. N. S. Wales, (2) 4 - via The Interim Register of Marine and Nonmarine Genera

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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