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Permembiidae ✝

Photos

Photo Gallery

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	8344373	https://www.gbif.org/species/8344373
PaleoBioDB ID (PBDB)	176244	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=176244

Rank

family

Taxonomy (GBIF)

Life : Animalia : Arthropoda : Insecta : Grylloblattodea : Permembiidae

Taxonomy (PBDB)

Life : Animalia : Arthropoda : Insecta : Permembiidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
family	Permembiidae	Tillyard 1937

Scientific Name

Permembiidae

Name Published In

Tillyard. 1937. Kansas Permian insects. Part 17. The order Megasecoptera and additions to the Paleodictyoptera, Odonata, Protoperlaria, Copeognata and Neuroptera. The American Journal of Science (Amer. Jour. Sci.) 33:81-110

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Permembiidae	family	belongs to Miomoptera	stated with evidence	Riek, 1973
Permembiidae	family	belongs to Protoperlina	stated with evidence	Storozhenko, 1997
Permembiidae	family	belongs to Hypoperlida	stated with evidence	Rasnitsyn and Aristov, 2013
Permembiidae	family	belongs to Neoptera	stated with evidence	Prokop et al., 2017

Status (PBDB)

extinct

Taxon Size (PBDB)

First Recorded Appearance

290 - 279 Ma
Permian

Last Recorded Appearance

260 - 254 Ma
Permian

Type Specimen

†Permembiidae Tillyard, 1937

Designated By

Type Information Source

Grylloblattodea Species File

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

R. J. Tillyard. 1937. Kansas Permian Insects, Part 17: The Order Megasecoptera and Additions to the Palaeodictyoptera, Odonata, Protoperlaria, Copeognatha, and Neuroptera. American Journal of Science, Series 5 33(194):81-110

Fossil Distribution

Subtaxa

Name	Status	Fossil Occurrences	Oldest	Youngest
Letopalopterinae ✝ subfamily	listed (PBDB)	8	279 Ma Permian	260 Ma Permian
Issapaloptera ✝ genus	accepted (GBIF) listed (PBDB)	1	260 Ma Permian	260 Ma Permian
Kungurmica ✝ genus	accepted (GBIF) listed (PBDB)	1	279 Ma Permian	279 Ma Permian
Letopaloptera ✝ genus	accepted (GBIF) listed (PBDB)	3	272 Ma Permian	272 Ma Permian
Neembia ✝ genus	accepted (GBIF) listed (PBDB)	1	260 Ma Permian	260 Ma Permian
Parasheimia ✝ genus	accepted (GBIF) listed (PBDB)	3	279 Ma Permian	272 Ma Permian
Permembia ✝ genus	accepted (GBIF) listed (PBDB)	2	290 Ma Permian	290 Ma Permian
Permindigena ✝ genus	accepted (GBIF) listed (PBDB)	3	279 Ma Permian	279 Ma Permian
Pseudosheimia ✝ genus	accepted (GBIF) listed (PBDB)	2	279 Ma Permian	272 Ma Permian
Sheimia ✝ genus	accepted (GBIF) listed (PBDB)	2	279 Ma Permian	272 Ma Permian
Soyanembia ✝ genus	accepted (GBIF) listed (PBDB)	1	272 Ma Permian	272 Ma Permian
Soyania ✝ genus	accepted (GBIF) listed (PBDB)	2	272 Ma Permian	272 Ma Permian
Visherifera ✝ genus	accepted (GBIF) listed (PBDB)	2	279 Ma Permian	279 Ma Permian

Synonymy List

Year	Name and Author
1937	Permembiidae Tillyard p. 92
1973	Permembiidae Riek p. 518
1997	Permembiidae Storozhenko p. 9
1998	Permembiidae Storozhenko p. 45
2004	Permembiidae Aristov
2009	Permembiidae Aristov p. 178
2013	Permembiidae Rasnitsyn and Aristov p. 695
2017	Permembiidae Prokop et al. p. 16

References

Tillyard R. J. (1937) Kansas Permian Insects, Part 17: The Order Megasecoptera and Additions to the Palaeodictyoptera, Odonata, Protoperlaria, Copeognatha, and Neuroptera, American Journal of Science, Series 5 33 194, 81-110 doi:10.2475/ajs.s5-33.194.81

Riek E. F. (1973) Fossil insects from the Upper Permian of Natal, South Africa, Annals of the Natal Museum 21 3, 513-532

Storozhenko S. Y. (1997) Classification of the Order Grylloblattida (Insecta), with description of new taxa, Far Eastern Entomologist 42, 1-20

Storozhenko S. Y. (1998) , Sistematika, Filogeniya i Evolyutsiya Grilloblattidovykh Nasekomykh (Insecta: Grylloblattida), 1-207

Aristov D. S. (2004) The Fauna of Grylloblattid Insects (Grylloblattida) of the Lower Permian Locality of Tshekarda, Paleontological Journal 38, S80-S145

Aristov D. S. (2009) A New Family of the Order Grylloblattida (Insecta) from the Middle Permian of Russia, Paleontological Journal 43 2, 178-182

Rasnitsyn A. P., Aristov D. S. (2013) New fossil insects (Insecta: Caloneurida, Hypoperlida, Palaeomanteida, Jurinida) from the Middle and Upper Permian of European Russia, Paleontological Journal 47, 678-704

Prokop J., Pecharová M., et al (2017) Redefining the extinct orders Miomoptera and Hypoperlida as stem acercarian insects, BMC Evolutionary Biology 17 205, 1-20

American Journal of Science Series 5 33 (194): -. [81-110] - via The Interim Register of Marine and Nonmarine Genera

Aristov & Rasnitsyn, 2008 - via The Interim Register of Marine and Nonmarine Genera

Aristov. 2004. The Fauna of Grylloblattid Insects (Grylloblattida) of the Lower Permian Locality of Tshekarda. Paleontological Journal 38(Suppl 2):S88 - via Grylloblattodea Species File

Martynova. 1962. In Rodendorf [Ed.]. Osnovy paleontologii. Chlenistonogie, trakheinye i khelitserovye. Order Raphidioptera. Academy of Science USSR Publ. (Acad. Sci. USSR Publ.) 270-272 - via Grylloblattodea Species File

Storozhenko. 1997. Classification of order Grylloblattida (Insecta), with description of new taxa. Far Eastern Entomologist 42:9 - via Grylloblattodea Species File

Storozhenko. 1997. System and phylogeny of the order Grylloblattida (Insecta). Chteniya Pamyati Alekseya Ivanovicha Kurentsova 7:25 - via Grylloblattodea Species File

Carpenter. 1976. The Lower Permian insects of Kansas. Part 12. Protorthoptera (continued), Neuroptera, aditional Paleodictyoptera, and families of uncertain position. Psyche 83(3-4):336-378 - via Grylloblattodea Species File

Storozhenko. 2002. In Rasnitsyn. Infraclass Gryllones. In Rasnitsyn & Quicke [Ed.]. Order Grylloblattida Walker, 1914. History of Insects, Kluwer Academic Publishers 278-283 - via Grylloblattodea Species File

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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