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Nemestrinidae

Description

Nemestrinidae, or tangle-veined flies is a family of flies in the superfamily Nemestrinoidea, closely related to Acroceridae. The family is small but distributed worldwide, with about 300 species in 34 genera. Larvae are endoparasitoids of either grasshoppers (Trichopsideinae) or scarab beetles (Hirmoneurinae). Some are considered important in the control of grasshopper populations. Adults are often observed on flowers.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)5569https://www.gbif.org/species/5569
PaleoBioDB ID (PBDB)138873https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=138873
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Diptera : Nemestrinidae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-Holometabola
-Aparaglossata
-Mecopterida
superorderAntliophora
orderDipteraLinnaeus 1758
-NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
-Homeodactyla
superfamilyNemestrinoideaMacquart 1834
familyNemestrinidaeMacquart 1834
Common Nametangle-veined fly
Scientific NameNemestrinidae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Nemestrinidaefamilybelongs to Brachycerastated without evidenceCarpenter, 1992
Nemestrinidaefamilybelongs to Nemestrinoideastated with evidenceMostovski and Martínez-Delclòs, 2000
Status (PBDB)extant
Taxon Size (PBDB)51
Extant Size (PBDB)9 (18%)
First Recorded Appearance166 - 157 Ma
Jurassic
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)F. M. Carpenter. 1992. Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4:1-655
Common Name(s) Tangle-veined Flies, Tangle-veined Fly
Wikipediahttps://en.wikipedia.org/wiki/Nemestrinidae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Archinemestriinae
subfamily
listed (PBDB)19166 Ma
Middle Jurassic
99.6 Ma
Late/Upper Cretaceous
Nemestrininae
subfamily
listed (PBDB)3130 Ma
Early/Lower Cretaceous
0 Ma
Extant
Aenigmestrinus
genus
accepted (GBIF)
listed (PBDB)
2166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Ahirmoneura
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Archinemestrius
genus
accepted (GBIF)
listed (PBDB)
6166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Atriadops
genus
accepted (GBIF)No associated record in PBDB
Ceyloniola
genus
accepted (GBIF)No associated record in PBDB
Cyclopsidea
genus
accepted (GBIF)No associated record in PBDB
Eohirmoneura
genus
accepted (GBIF)No associated record in PBDB
Fallenia
genus
accepted (GBIF)No associated record in PBDB
Florinemestrius
genus
accepted (GBIF)
listed (PBDB)
1125 Ma
Early/Lower Cretaceous
125 Ma
Early/Lower Cretaceous
Hirmoneura
genus
accepted (GBIF)
listed (PBDB)
12166 Ma
Middle Jurassic
0 Ma
Extant
Hirmoneurinae
genus
accepted (GBIF)
listed (PBDB)
13166 Ma
Middle Jurassic
0 Ma
Extant
Hyrmophlaeba
genus
accepted (GBIF)No associated record in PBDB
Moegistorhynchus
genus
accepted (GBIF)No associated record in PBDB
Nemestrina
genus
listed (PBDB)0 Ma
Extant
Nemestrinus
genus
accepted (GBIF)
listed (PBDB)
1125 Ma
Early/Lower Cretaceous
0 Ma
Extant
Neorhynchocephalus
genus
accepted (GBIF)
listed (PBDB)
737.2 Ma
Eocene
0 Ma
Extant
Nycterimorpha
genus
accepted (GBIF)No associated record in PBDB
Nycterimyia
genus
accepted (GBIF)No associated record in PBDB
Palembolus
genus
accepted (GBIF)No associated record in PBDB
Prohirmoneura
genus
accepted (GBIF)
listed (PBDB)
1151 Ma
Late Jurassic
151 Ma
Late Jurassic
Prosoeca
genus
accepted (GBIF)
listed (PBDB)
2130 Ma
Early/Lower Cretaceous
0 Ma
Extant
Protonemestrius
genus
accepted (GBIF)
listed (PBDB)
10166 Ma
Middle Jurassic
125 Ma
Early/Lower Cretaceous
Rhagionemestrius
genus
accepted (GBIF)
listed (PBDB)
1166 Ma
Middle Jurassic
166 Ma
Middle Jurassic
Sinonemestrius
genus
accepted (GBIF)
listed (PBDB)
2136 Ma
Early/Lower Cretaceous
125 Ma
Early/Lower Cretaceous
Stenobasipteron
genus
accepted (GBIF)No associated record in PBDB
Stenopteromyia
genus
accepted (GBIF)No associated record in PBDB
Trichophthalma
genus
accepted (GBIF)No associated record in PBDB
Trichopsidea
genus
accepted (GBIF)No associated record in PBDB
Hyrmaphloeba surcana
species
accepted (GBIF)No associated record in PBDB
Rhynchocephala subnitens
species
accepted (GBIF)No associated record in PBDB

Synonymy List

YearName and Author
1834Nemestrinidae Macquart p. 370
1992Nemestrinidae Carpenter
2000Nemestrinidae Mostovski and Martínez-Delclòs p. 142
2016Nemestrinidae Grimaldi p. 49
2018Nemestrinidae Shin et al.

References

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655
Mostovski M. B., Martínez-Delclòs X. (2000) New Nemestrinoidea (Diptera: Brachycera) from the Upper Jurassic - Lower Cretaceous of Eurasia, taxonomy, and palaeobiology, Entomological Problems 31 2, 137-148
Grimaldi D. A. (2016) Diverse orthorrhaphan flies (Insecta: Diptera: Brachycera) in amber from the Cretaceous of Myanmar: Brachycera in Cretaceous amber, part VII, Bulletin of the American Museum of Natural History 408, 1-131
Shin S., Bayless K. M., et al (2018) Taxon sampling to address an ancient rapid radiation: a supermatrix phylogeny of early brachyceran flies (Diptera), Systematic Entomology doi:10.1111/syen.12275
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
Wood, D. M.,Teskey, H. J.,Peterson, B. V.,McAlpine, J. F.,Vockeroth, J. R.,Shewell, G. E. (1981) Chapter 44. Nemestrinidae - via The Diptera Site
Hardy, G. H. (1924) Australian Nemestrinidae (Diptera) - via The Diptera Site
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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