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Cassidinae

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Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	Not linked to a GBIF record.
PaleoBioDB ID (PBDB)	207310	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=207310

Rank

subfamily (PBDB)

Taxonomy (PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : Cassidinae

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Polyphaga	Emery 1886
infraorder	Cucujiformia	Lameere 1938
-	Phytophaga
superfamily	Chrysomeloidea	Latreille 1802
family	Chrysomelidae	Latreille 1802
subfamily	Cassidinae	Gyllenhal 1813

Common Name

tortoise beetle

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Cassidinae	subfamily	belongs to Chrysomelidae	stated with evidence	Haupt, 1950

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

19 (30%)

First Recorded Appearance

56.0 - 47.8 Ma
Eocene

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

P. Bouchard, Y. Bousquet, and A. E. Davies, M. A. Alonso-Zarazaga, J. F. Lawrence, C. H. C. Lyal, A. F. Newton, C. A. M. Reid, M. Schmitt, A. lipiski, A. B. T. Smith. 2011. Family-group names in Coleoptera (Insecta). ZooKeys 88:1-972

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Anisoderini tribe	listed (PBDB)		1	38.0 Ma Eocene	0 Ma Extant
Cassidini genus	doubtful (GBIF) listed (PBDB)		16	56.0 Ma Paleocene	0 Ma Extant
Chalepini tribe	listed (PBDB)		1	37.2 Ma Eocene	0 Ma Extant
Gonophorini tribe	listed (PBDB)		3	12.7 Ma Miocene	0 Ma Extant
Hispini tribe	listed (PBDB)	Leaf-mining Beetle	2	23.0 Ma Miocene	0 Ma Extant
Imatidiini tribe	listed (PBDB)		1	50.3 Ma Eocene	0 Ma Extant
Mesomphaliini tribe	listed (PBDB)		1	47.8 Ma Eocene	0 Ma Extant
Notosacanthini tribe	listed (PBDB)		1	46.2 Ma Eocene	0 Ma Extant
Oposispini ✝ tribe	listed (PBDB)		3	38.0 Ma Eocene	38.0 Ma Eocene
Uroplatini tribe	listed (PBDB)				0 Ma Extant
Delocrania genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	0 Ma Extant
Echinocallispa ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Electrolema ✝ genus	accepted (GBIF) listed (PBDB)		1	38.0 Ma Eocene	38.0 Ma Eocene
Oligocassida ✝ genus	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Paracassida ✝ genus	accepted (GBIF) listed (PBDB)		4	47.8 Ma Eocene	47.8 Ma Eocene
Sceloenopla genus	doubtful (GBIF) listed (PBDB)		3	20.4 Ma Miocene	0 Ma Extant

Synonymy List

Year	Name and Author
1813	Cassidinae Gyllenhal
1813	Hispinae Gyllenhal
1950	Cassidinae Haupt p. 153
1956	Cassidinae Haupt p. 74
1994	Hispidae Zhang et al. p. 133
1995	Hispinae Lawrence and Newton, Jr. p. 787
1996	Hispinae Santiago-Blay et al. p. 421

Year	Name and Author
1996	Cassidinae Santiago-Blay et al. p. 422
2003	Hispinae Wappler p. 109
2003	Cassidinae Wappler p. 110
2011	Cassidinae Bouchard et al. p. 510
2015	Cassidinae Nadein p. 118
2016	Cassidinae Webster

References

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Haupt H. (1950) Die Käfer (Coleoptera) aus der eozänen Braunkohle des Geiseltales, Geologica 6, 1-168

Haupt H. (1956) Beitrag zur Kenntnis der eozänen Arthopodenfauna des Geiseltales, Nova Acta Leopoldina 18 128, 1-89

Zhang J., Sun B., et al (1994) , Miocene insects and spiders from Shanwang, Shandong, 1-298

Lawrence J. F., Newton, Jr. A. F. (1995) Families and subfamilies of Coleoptera, Biology, Phylogeny and Classification of Coleoptera: Papers Celebrating the 80th Birthday of Roy A Crowson, 779-1006

Santiago-Blay J. A., Poinar G. O., et al (1996) Dominican and Mexican amber chrysomelids, with descriptions of two new species, Chrysomelid Biology, vol. 1: The Classification, Phylogeny and Genetics, 413-424

Wappler T. (2003) Systematik, Phylogenie, Taphonomie und Paläoökologie der Insekten aus dem Mittel-Eozän des Eckfelder Maares, Vulkaneifel, Clausthaler Geowissenschaften 2, 1-241

Nadein K. S. (2015) Taxonomic names, in New late Eocene Chrysomelidae (Insecta: Coleoptera) from Baltic, Rovno and Danish ambers, Papers in Palaeontology 2, 117-137

Webster R. P. (2016) Checklist of the Coleoptera of New Brunswick, Canada, ZooKeys 573, 387-512

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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