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Carbonera

Formation	Carbonera
Age:	23.0 - 11.6 Ma Miocene
Interval	Berriasian
Lithology	shale/sandstone, sandstone, "shale", "siliciclastic", sandstone/siltstone
Number of Collections	14
Number of Occurrences	171

Recorded Sample Locations

Location	Region	Stratigraphic Name
A3-Eastern Durango	General Francisco Murguía Municipality, Zacatecas, Mexico	Carbonera
18-Coahuila	Cuatro Ciénegas Municipality, Coahuila, Mexico	Carbonera
Regadera UFP 53	Norte de Santander Department, Colombia	Carbonera
Hannatoma A: Carbonera Fm: El Cerrito	Santander Department, Colombia	Carbonera
Hannatoma C: Carbonera Fm: Quebrada Seca	Cesar Department, Colombia	Carbonera
Gaz 39.Gazaunta River. Carbonera Formation	Cundinamarca Department, Colombia	Carbonera

Associated Units

Stratigraphic Name	Age	Lithology	Occurrence Records
Cabonera	145 - 140 Ma Early/Lower Cretaceous	sandstone/"shale"	1
Cupido	129 - 125 Ma Early/Lower Cretaceous		1
La Casita	151 - 145 Ma Late Jurassic	not reported/siltstone, siltstone, "limestone"	24
Mirador	48.6 - 37.2 Ma Eocene	sandstone/mudstone, mudstone, "siliciclastic", sandstone, sandstone/"siliciclastic", sandstone/"shale"	701
Taraises	133 - 129 Ma Early/Lower Cretaceous	marl, "limestone"	39

Recorded Fossils

Accepted Name	Type	Hierarchy	Age
Mauritiidites franciscoi species	7	Plantae : Tracheophyta : Liliopsida : Arecales : Arecaceae : Mauritiidites : *Mauritiidites franciscoi*	48.6 - 37.2 Ma Eocene
Longapertites proxapertitoides species	7	Plantae : Tracheophyta : Liliopsida : Arecales : Longapertites : *Longapertites proxapertitoides*	48.6 - 37.2 Ma Eocene
Monoporopollenites annulatus species	7	Plantae : Tracheophyta : Liliopsida : Poales : Poaceae : Monoporopollenites : *Monoporopollenites annulatus*	48.6 - 37.2 Ma Eocene
Tetracolporopollenites sp. genus	6	*Tetracolporopollenites*	48.6 - 37.2 Ma Eocene
Verrutricolporites sp. genus	6	*Verrutricolporites*	48.6 - 37.2 Ma Eocene
Cyclusphaera sp. genus	6	Plantae : Tracheophyta : Pinopsida : Pinales : Araucariaceae : *Cyclusphaera*	48.6 - 37.2 Ma Eocene
Lanagiopollis crassa species	7	Plantae : Tracheophyta : Magnoliopsida : Ericales : Lanagiopollis : *Lanagiopollis crassa*	48.6 - 37.2 Ma Eocene
Spirosyncolpites spiralis species	7	Plantae : Tracheophyta : Magnoliopsida : Fabales : Fabaceae : Spirosyncolpites : *Spirosyncolpites spiralis*	48.6 - 37.2 Ma Eocene
Nothofagidites sp. genus	6	Plantae : Tracheophyta : Magnoliopsida : Fagales : *Nothofagidites*	48.6 - 37.2 Ma Eocene
Echiperiporites estelae species	7	Plantae : Tracheophyta : Magnoliopsida : Malvales : Echiperiporites : *Echiperiporites estelae*	48.6 - 37.2 Ma Eocene
Baculamonocolpites hammenii species	7	Plantae : Tracheophyta : Magnoliopsida : Baculamonocolpites : *Baculamonocolpites hammenii*	48.6 - 37.2 Ma Eocene
Clavatricolpites densiclavatus species	7	Plantae : Tracheophyta : Magnoliopsida : Clavatricolpites : *Clavatricolpites densiclavatus*	48.6 - 37.2 Ma Eocene
Echitetracolpites sp. genus	6	Plantae : Tracheophyta : Magnoliopsida : *Echitetracolpites*	48.6 - 37.2 Ma Eocene
Foveotricolporites rugulatus species	7	Plantae : Tracheophyta : Magnoliopsida : Foveotricolporites : *Foveotricolporites rugulatus*	48.6 - 37.2 Ma Eocene
Perisyncolporites pokornyi species	7	Plantae : Tracheophyta : Magnoliopsida : Malpighiales : Malpighiaceae : Perisyncolporites : *Perisyncolporites pokornyi*	48.6 - 37.2 Ma Eocene
Echitriporites trianguliformis species	7	Plantae : Spermatophyta : Magnoliopsida : Proteales : Proteaceae : Echitriporites : *Echitriporites trianguliformis*	48.6 - 37.2 Ma Eocene
Rhynchonella durangensis species	7	Animalia : Brachiopoda : Rhynchonellata : Rhynchonellida : Rhynchonellidae : Rhynchonella : *Rhynchonella durangensis*	145 - 139.8 Ma Early/Lower Cretaceous
Antiptychina muellerriedi species	7	Animalia : Brachiopoda : Rhynchonellata : Terebratulida : Zeilleriidae : Antiptychina : *Antiptychina muellerriedi*	145 - 139.8 Ma Early/Lower Cretaceous
Paretallonia hispaniensis species	7	Animalia : Cnidaria : Anthozoa : Scleractinia : Acroporidae : Paretallonia : *Paretallonia hispaniensis*	145 - 139.8 Ma Early/Lower Cretaceous
Arginella puntabravoensis species	7	Animalia : Mollusca : Bivalvia : Arcida : Arcidae : Arginella : *Arginella puntabravoensis*	37.2 - 33.9 Ma Paleogene
Arginella samanensis species	7	Animalia : Mollusca : Bivalvia : Arcida : Arcidae : Arginella : *Arginella samanensis*	37.2 - 33.9 Ma Paleogene
Cucullaea sp. genus	6	Animalia : Mollusca : Bivalvia : Arcida : Cucullaeidae : *Cucullaea*	145 - 139.8 Ma Early/Lower Cretaceous
Cucullaea striatiumbonata species	7	Animalia : Mollusca : Bivalvia : Arcida : Cucullaeidae : Cucullaea : *Cucullaea striatiumbonata*	145 - 139.8 Ma Early/Lower Cretaceous
Tagelus bolivarensis species	7	Animalia : Mollusca : Bivalvia : Cardiida : Solecurtidae : Tagelus : *Tagelus bolivarensis*	37.2 - 33.9 Ma Paleogene
Macoma sp. genus	6	Animalia : Mollusca : Bivalvia : Cardiida : Tellinidae : *Macoma*	37.2 - 33.9 Ma Paleogene
Tellina sp. genus	6	Animalia : Mollusca : Bivalvia : Cardiida : Tellinidae : *Tellina*	37.2 - 33.9 Ma Paleogene
Astarte durangensis species	7	Animalia : Mollusca : Bivalvia : Carditida : Astartidae : Astarte : *Astarte durangensis*	145 - 139.8 Ma Early/Lower Cretaceous
Phacoides sp. genus	6	Animalia : Mollusca : Bivalvia : Lucinida : Lucinidae : *Phacoides*	37.2 - 33.9 Ma Paleogene
Exogyra reedi species	7	Animalia : Mollusca : Bivalvia : Ostreida : Gryphaeidae : Exogyra : *Exogyra reedi*	145 - 139.8 Ma Early/Lower Cretaceous
Exogyra tamaulipana species	7	Animalia : Mollusca : Bivalvia : Ostreida : Gryphaeidae : Exogyra : *Exogyra tamaulipana*	145 - 139.8 Ma Early/Lower Cretaceous
Gryphaea sp. genus	6	Animalia : Mollusca : Bivalvia : Ostreida : Gryphaeidae : *Gryphaea*	145 - 139.8 Ma Early/Lower Cretaceous
Ostrea sp. genus	6	Animalia : Mollusca : Bivalvia : Ostreida : Ostreidae : *Ostrea*	37.2 - 33.9 Ma Paleogene
Ostrea (Alectryonia) sp. subgenus	6.3	Animalia : Mollusca : Bivalvia : Ostreida : Ostreidae : Ostrea : *Ostrea (Alectryonia)*	145 - 139.8 Ma Early/Lower Cretaceous
Pinna sp. genus	6	Animalia : Mollusca : Bivalvia : Ostreida : Pinnidae : *Pinna*	145 - 139.8 Ma Early/Lower Cretaceous
Anomia sp. genus	6	Animalia : Mollusca : Bivalvia : Pectinida : Anomiidae : *Anomia*	37.2 - 33.9 Ma Paleogene
Carolia sp. genus	6	Animalia : Mollusca : Bivalvia : Pectinida : Anomiidae : *Carolia*	37.2 - 33.9 Ma Paleogene
Camptonectes sp. genus	6	Animalia : Mollusca : Bivalvia : Pectinida : Pectinidae : *Camptonectes*	145 - 139.8 Ma Early/Lower Cretaceous
Neithea (Neithea) atava species	7	Animalia : Mollusca : Bivalvia : Pectinida : Pectinidae : Neithea : *Neithea (Neithea) atava*	145 - 139.8 Ma Early/Lower Cretaceous
Pecten sp. genus	6	Animalia : Mollusca : Bivalvia : Pectinida : Pectinidae : *Pecten*	145 - 139.8 Ma Early/Lower Cretaceous
Plicatula sp. genus	6	Animalia : Mollusca : Bivalvia : Pectinida : Plicatulidae : *Plicatula*	145 - 139.8 Ma Early/Lower Cretaceous
Plicatula azulensis species	7	Animalia : Mollusca : Bivalvia : Pectinida : Plicatulidae : Plicatula : *Plicatula azulensis*	145 - 139.8 Ma Early/Lower Cretaceous
Plicatula pulchra species	7	Animalia : Mollusca : Bivalvia : Pectinida : Plicatulidae : Plicatula : *Plicatula pulchra*	145 - 139.8 Ma Early/Lower Cretaceous
Mya sp. genus	6	Animalia : Mollusca : Bivalvia : Myida : Myidae : *Mya*	37.2 - 33.9 Ma Paleogene
Corbicula sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Cyrenidae : *Corbicula*	37.2 - 33.9 Ma Paleogene
Cyrena sp. genus	6	Animalia : Mollusca : Bivalvia : Veneroidei : Cyrenidae : *Cyrena*	37.2 - 33.9 Ma Paleogene
Mactra sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Mactridae : *Mactra*	37.2 - 33.9 Ma Paleogene
Trapezium sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Trapezidae : *Trapezium*	145 - 139.8 Ma Early/Lower Cretaceous
Trapezium subtriangulum species	7	Animalia : Mollusca : Bivalvia : Venerida : Trapezidae : Trapezium : *Trapezium subtriangulum*	145 - 139.8 Ma Early/Lower Cretaceous
Agriopoma sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Veneridae : *Agriopoma*	37.2 - 33.9 Ma Paleogene
Clementia (Egesta) peruviana species	7	Animalia : Mollusca : Bivalvia : Veneroidei : Veneridae : Clementia : *Clementia (Egesta) peruviana*	37.2 - 33.9 Ma Paleogene
Cyclorisma sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Veneridae : *Cyclorisma*	145 - 139.8 Ma Early/Lower Cretaceous
Pitar sp. genus	6	Animalia : Mollusca : Bivalvia : Venerida : Veneridae : *Pitar*	37.2 - 33.9 Ma Paleogene
Pitar (Pitarella) colombiana species	7	Animalia : Mollusca : Bivalvia : Veneroidei : Veneridae : Agriopoma : *Pitar (Pitarella) colombiana*	37.2 - 33.9 Ma Paleogene
Transennella bolivarensis species	7	Animalia : Mollusca : Bivalvia : Venerida : Veneridae : Transennella : *Transennella bolivarensis*	37.2 - 33.9 Ma Paleogene
Hemihoplites mexicanus species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Hemihoplitidae : Hemihoplites : *Hemihoplites mexicanus*	132.9 - 129.4 Ma Early/Lower Cretaceous
Lytoceras sp. genus	6	Animalia : Mollusca : Cephalopoda : Ammonitida : Lytoceratidae : *Lytoceras*	145 - 139.8 Ma Early/Lower Cretaceous
Berriasella sp. genus	6	Animalia : Mollusca : Cephalopoda : Ammonoidea : Neocomitidae : *Berriasella*	145 - 139.8 Ma Early/Lower Cretaceous
Distoloceras sp. genus	6	Animalia : Mollusca : Cephalopoda : Ammonoidea : Neocomitidae : *Distoloceras*	132.9 - 129.4 Ma Early/Lower Cretaceous
Thurmanniceras sp. genus	6	Animalia : Mollusca : Cephalopoda : Ammonitida : Neocomitidae : *Thurmanniceras*	145 - 139.8 Ma Early/Lower Cretaceous
Thurmannites miquihuanensis species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Neocomitidae : Thurmanniceras : *Thurmannites miquihuanensis*	145 - 139.8 Ma Early/Lower Cretaceous
Thurmannites novihispanicus species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Neocomitidae : Thurmanniceras : *Thurmannites novihispanicus*	145 - 139.8 Ma Early/Lower Cretaceous
Maderia casitensis species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Olcostephanidae : Maderia : *Maderia casitensis*	132.9 - 129.4 Ma Early/Lower Cretaceous
Mexicanoceras laticostatum species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Olcostephanidae : Mexicanoceras : *Mexicanoceras laticostatum*	132.9 - 129.4 Ma Early/Lower Cretaceous
Mexicanoceras multicostatum species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Olcostephanidae : Mexicanoceras : *Mexicanoceras multicostatum*	132.9 - 129.4 Ma Early/Lower Cretaceous
Blanforidceras sp. genus	6	Animalia : Mollusca : Cephalopoda : Ammonoidea : *Blanforidceras*	145 - 139.8 Ma Early/Lower Cretaceous
Phylloceras serum species	7	Animalia : Mollusca : Cephalopoda : Ammonoidea : Phylloceratidae : Phylloceras : *Phylloceras serum*	132.9 - 129.4 Ma Early/Lower Cretaceous
Peruficus lagunitensis species	7	Animalia : Mollusca : Gastropoda : Neogastropoda : Fasciolariidae : Glyptostyla : *Peruficus lagunitensis*	37.2 - 33.9 Ma Paleogene
Mazzalina sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Fasciolariidae : *Mazzalina*	37.2 - 33.9 Ma Paleogene
Cornulina sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Melongenidae : *Cornulina*	37.2 - 33.9 Ma Paleogene
Melongena sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Melongenidae : *Melongena*	37.2 - 33.9 Ma Paleogene
Agaronia harrisi species	7	Animalia : Mollusca : Gastropoda : Neogastropoda : Olividae : Agaronia : *Agaronia harrisi*	37.2 - 33.9 Ma Paleogene
Oliva sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Olividae : *Oliva*	37.2 - 33.9 Ma Paleogene
Terebra sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Terebridae : *Terebra*	37.2 - 33.9 Ma Paleogene
Lyria sp. genus	6	Animalia : Mollusca : Gastropoda : Neogastropoda : Volutidae : *Lyria*	37.2 - 33.9 Ma Paleogene
Crommium palmerae species	7	Animalia : Mollusca : Gastropoda : Architaenioglossa : Ampullinidae : Crommium : *Crommium palmerae*	37.2 - 33.9 Ma Paleogene
Natica sp. genus	6	Animalia : Mollusca : Gastropoda : Littorinimorpha : Naticidae : *Natica*	37.2 - 33.9 Ma Paleogene
Neverita sp. genus	6	Animalia : Mollusca : Gastropoda : Littorinimorpha : Naticidae : *Neverita*	37.2 - 33.9 Ma Paleogene
Neverita (Glossaulax) bolivarensis species	7	Animalia : Mollusca : Gastropoda : Naticidae : Neverita : *Neverita (Glossaulax) bolivarensis*	37.2 - 33.9 Ma Paleogene
Polinices sp. genus	6	Animalia : Mollusca : Gastropoda : Naticidae : *Polinices*	37.2 - 33.9 Ma Paleogene
Rimella sp. genus	6	Animalia : Mollusca : Gastropoda : Strombidae : *Rimella*	37.2 - 33.9 Ma Paleogene
Cerithiella sp. genus	6	Animalia : Mollusca : Gastropoda : Newtoniellidae : *Cerithiella*	37.2 - 33.9 Ma Paleogene
Vermetus cornejoi species	7	Animalia : Mollusca : Gastropoda : Littorinimorpha : Vermetidae : Vermetus : *Vermetus cornejoi*	145 - 139.8 Ma Early/Lower Cretaceous
Pleurotomaria sp. genus	6	Animalia : Mollusca : Gastropoda : Trochida : Microdomatidae : *Pleurotomaria*	145 - 139.8 Ma Early/Lower Cretaceous
Cerithium (Perucerithium) sp. subgenus	6.3	Animalia : Mollusca : Gastropoda : Sorbeoconcha : Cerithiidae : Cerithium : *Cerithium (Perucerithium)*	37.2 - 33.9 Ma Paleogene
Diastoma sp. genus	6	Animalia : Mollusca : Gastropoda : Diastomatidae : *Diastoma*	37.2 - 33.9 Ma Paleogene
Faunus (Pseudofaunus) sp. subgenus	6.3	Animalia : Mollusca : Gastropoda : Sorbeoconcha : Pachychilidae : Faunus : *Faunus (Pseudofaunus)*	37.2 - 33.9 Ma Paleogene
Melanatria sp. genus	6	Animalia : Mollusca : Gastropoda : Pachychilidae : *Melanatria*	37.2 - 33.9 Ma Paleogene
Hannatoma emendorferi species	7	Animalia : Mollusca : Gastropoda : Thiaridae : Hannatoma : *Hannatoma emendorferi*	37.2 - 33.9 Ma Paleogene
Harrisianella sp. genus	6	Animalia : Mollusca : Gastropoda : Potamididae : *Harrisianella*	37.2 - 33.9 Ma Paleogene
Harrisianella peruviana species	7	Animalia : Mollusca : Gastropoda : Potamididae : Harrisianella : *Harrisianella peruviana*	37.2 - 33.9 Ma Paleogene
Lagunitus sp. genus	6	Animalia : Mollusca : Gastropoda : Potamididae : *Lagunitus*	37.2 - 33.9 Ma Paleogene
Potamides sp. genus	6	Animalia : Mollusca : Gastropoda : Sorbeoconcha : Potamididae : *Potamides*	37.2 - 33.9 Ma Paleogene
Telescopium sp. genus	6	Animalia : Mollusca : Gastropoda : Potamididae : *Telescopium*	37.2 - 33.9 Ma Paleogene
Turritella sp. genus	6	Animalia : Mollusca : Gastropoda : Turritellidae : *Turritella*	37.2 - 33.9 Ma Paleogene
Turritella chira species	7	Animalia : Mollusca : Gastropoda : Turritellidae : Turritella : *Turritella chira*	37.2 - 33.9 Ma Paleogene
Turritella samanensis species	7	Animalia : Mollusca : Gastropoda : Turritellidae : Turritella : *Turritella samanensis*	37.2 - 33.9 Ma Paleogene
Botryococcus sp. genus	6	Plantae : Chlorophyta : Trebouxiophyceae : Trebouxiales : Botryococcaceae : *Botryococcus*	23.03 - 11.608 Ma Miocene
Pediastrum sp. genus	6	Plantae : Chlorophyta : Chlorophyceae : Sphaeropleales : Hydrodictyaceae : *Pediastrum*	23.03 - 11.608 Ma Miocene
Proxapertites operculatus species	7	Plantae : Magnoliophyta : Proxapertites : *Proxapertites operculatus*	48.6 - 37.2 Ma Eocene
Psilamonocolpites medius species	7	Plantae : Magnoliophyta : Psilamonocolpites : *Psilamonocolpites medius*	48.6 - 37.2 Ma Eocene
Retistephanocolpites fossulatus species	7	Plantae : Magnoliophyta : Retistephanocolpites : *Retistephanocolpites fossulatus*	48.6 - 37.2 Ma Eocene
Retistephanoporites angelicus species	7	Plantae : Magnoliophyta : Retistephanoporites : *Retistephanoporites angelicus*	48.6 - 37.2 Ma Eocene
Retisyncolporites angularis species	7	Plantae : Magnoliophyta : Retisyncolporites : *Retisyncolporites angularis*	48.6 - 37.2 Ma Eocene
Retitrescolpites irregularis species	7	Plantae : Magnoliophyta : Retitrescolpites : *Retitrescolpites irregularis*	48.6 - 37.2 Ma Eocene
Scabratriporites sp. genus	6	Plantae : Magnoliophyta : *Scabratriporites*	48.6 - 37.2 Ma Eocene
Psilatriletes sp. genus	6	Plantae : Pteridophyta : *Psilatriletes*	23.03 - 11.608 Ma Miocene
Echinatisporis brevispinosus species	7	Plantae : Pteridophyta : Pteridopsida : Echinatisporis : *Echinatisporis brevispinosus*	48.6 - 37.2 Ma Eocene
Laevigatosporites tibuensis species	7	Plantae : Pteridophyta : Pteridopsida : Laevigatosporites : *Laevigatosporites tibuensis*	48.6 - 37.2 Ma Eocene
Polypodiisporites sp. genus	6	Plantae : Tracheophyta : Polypodiopsida : Polypodiales : *Polypodiisporites*	48.6 - 37.2 Ma Eocene
Polypodiisporites inangahuensis species	7	Plantae : Tracheophyta : Polypodiopsida : Polypodiales : Polypodiisporites : *Polypodiisporites inangahuensis*	48.6 - 37.2 Ma Eocene
Cicatricosisporites dorogensis species	7	Plantae : Tracheophyta : Polypodiopsida : Schizaeales : Schizaeaceae : Cicatricosisporites : *Cicatricosisporites dorogensis*	48.6 - 37.2 Ma Eocene

References

Imlay R. W. (1940) Neocomian Faunas of Northern Mexico, Geological Society America Bulletin 51 1, 117-190

Durham J. W., Dusenbury, Jr A.N., et al (1949) The age of the Hannatoma mollusk fauna of South America; a symposium, Journal of Paleontology 23 2, 145-160

Jaramillo C. A., Dilcher D. L. (2001) Middle Paleogene palynology of Central Colombia, South America: A study of pollen and spores from tropical latitudes, Palaeontographica Abteilung B 258 4-6, 87-213

Gomez A., Jaramillo C., et al (2009) Huesser Horizon: A lake and marine incursion in Northwestern South America during the Early Miocene, Palaios 24, 199-210 doi:10.2110/palo.2007.p07-074r

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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