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Brule - upper Whitney, Nebraska, USA

Formation	Brule
Member	upper Whitney
Age:	30.8 - 20.4 Ma Cenozoic
Interval	Whitneyan
Number of Collections	21
Number of Occurrences	36

Recorded Sample Locations

Location	Region	Stratigraphic Name
Chimney Rock	Morrill County, Nebraska, USA	Brule - upper Whitney
Bald Butte	Sioux County, Nebraska, USA	Brule - upper Whitney
Lyman Beaver Locality	Scotts Bluff County, Nebraska, USA	Brule - Upper Whitney

Associated Units

Stratigraphic Name	Age	Lithology	Occurrence Records
Arikaree	20.4 - 5.33 Ma Miocene	siliciclastic	278
Ash Hollow	10.3 - 4.90 Ma Neogene	siliciclastic	231
Brown Siltstone Beds	30.8 - 20.4 Ma Cenozoic		2
Brule	33.3 - 30.8 Ma Oligocene	sandstone	162
Brule - Brown Siltstone	30.8 - 20.4 Ma Cenozoic		47
Brule - lower Orella	33.9 - 33.3 Ma Oligocene		213
Brule - lower Whitney	33.3 - 30.8 Ma Oligocene		4
Brule - Orella	33.9 - 33.3 Ma Oligocene		135
Brule - Orella D	33.9 - 33.3 Ma Oligocene		47
Brule - upper	30.8 - 20.4 Ma Cenozoic		5
Brule - Whitney	30.8 - 20.4 Ma Cenozoic		141
Brule - Whitney A	33.3 - 30.8 Ma Oligocene		16
Chadron	37.2 - 33.9 Ma Paleogene	claystone, "siliciclastic", "limestone", "volcaniclastic"	132
Chadron - Big Cottonwood Creek	37.2 - 33.9 Ma Paleogene	siltstone	174
Chadron - Peanut Peak	37.2 - 33.3 Ma Paleogene		85
Gering	30.8 - 26.3 Ma Oligocene		29
Gering - Gering A	30.8 - 26.3 Ma Oligocene		24
Gering - lower	30.8 - 26.3 Ma Oligocene		4
Gering - Unit B	30.8 - 26.3 Ma Oligocene		41
Gering - Unit C	30.8 - 26.3 Ma Oligocene		21
Gering - Unit D	30.8 - 26.3 Ma Oligocene		10
Harrison	24.8 - 20.4 Ma Cenozoic	sandstone	160
Monroe Creek	26.3 - 20.4 Ma Cenozoic		102
Olcott	16.0 - 13.6 Ma Miocene		109
Olcott - subcycle 1	20.4 - 16.0 Ma Miocene		23
Olcott - subcycle 2	20.4 - 16.0 Ma Miocene		44
Olcott - subcycle 3	20.4 - 16.0 Ma Miocene		65
Runningwater	20.4 - 16.0 Ma Miocene	sandstone, sandstone/conglomerate	315
Sheep Creek	20.4 - 16.0 Ma Miocene	sandstone/conglomerate	29
Sheep Creek - lower	20.4 - 16.0 Ma Miocene		40
Sheep Creek - middle	20.4 - 16.0 Ma Miocene	sandstone	96
Snake Creek	13.6 - 10.3 Ma Miocene		4
Snake Creek - Johnson	10.3 - 4.90 Ma Neogene		51
Snake Creek - Laucomer	13.6 - 10.3 Ma Miocene		43
Snake Creek - Murphy	13.6 - 10.3 Ma Miocene		3
Snake Creek - subcycle 3	20.4 - 16.0 Ma Miocene		3
Arikaree - Harrison	23.0 - 5.33 Ma Miocene		32
Arikaree - Harrison - Lower	24.8 - 20.4 Ma Cenozoic		47
Hemingford - Marsland	24.8 - 20.4 Ma Cenozoic	sandstone	107
Hemingford - Marsland - Basal	24.8 - 20.4 Ma Cenozoic	sandstone	44
White River - Brule	33.9 - 33.3 Ma Oligocene		25

Recorded Fossils

Accepted Name	Type	Hierarchy	Age
Paratylopus primaevus species	7	Animalia : Chordata : Mammalia : Artiodactyla : Camelidae : Paratylopus : *Paratylopus primaevus*	33.3 - 30.8 Ma Oligocene
Leptochoerus spectabilis species	7	Animalia : Chordata : Mammalia : Artiodactyla : Leptochoeridae : Leptochoerus : *Leptochoerus spectabilis*	33.3 - 30.8 Ma Oligocene
Leptauchenia decora species	7	Animalia : Chordata : Mammalia : Artiodactyla : Merycoidodontidae : Leptauchenia : *Leptauchenia decora*	33.3 - 30.8 Ma Oligocene
Leptauchenia major species	7	Animalia : Chordata : Mammalia : Artiodactyla : Merycoidodontidae : Leptauchenia : *Leptauchenia major*	33.3 - 30.8 Ma Oligocene
Merycoidodon (Otarohyus) major species	7	Animalia : Chordata : Mammalia : Artiodactyla : Merycoidodontidae : Merycoidodon : *Merycoidodon (Otarohyus) major*	33.3 - 30.8 Ma Oligocene
Mesoreodon minor species	7	Animalia : Chordata : Mammalia : Artiodactyla : Merycoidodontidae : Mesoreodon : *Mesoreodon minor*	33.3 - 30.8 Ma Oligocene
Cynodictis temnodon species	7	Animalia : Chordata : Mammalia : Carnivora : Canidae : Mesocyon : *Cynodictis temnodon*	33.3 - 30.8 Ma Oligocene
Hyracodon sp. genus	6	Animalia : Chordata : Mammalia : Perissodactyla : Hyracodontidae : *Hyracodon*	30.8 - 20.43 Ma Cenozoic
Hyracodon nebraskensis species	7	Animalia : Chordata : Mammalia : Perissodactyla : Hyracodontidae : Hyracodon : *Hyracodon nebraskensis*	33.3 - 30.8 Ma Oligocene
Campestrallomys siouxensis species	7	Animalia : Chordata : Mammalia : Rodentia : Allomyidae : Campestrallomys : *Campestrallomys siouxensis*	33.3 - 30.8 Ma Oligocene

References

Schultz C. B., Falkenbach C. H. (1954) Desmatochoerinae, a new subfamily of oreodonts, Bulletin of the American Museum of Natural History 105 2

Schultz C. B., Falkenbach C. H. (1968) The Phylogeny of the Oreodonts. Part 1: Merycoidodontinae, Eporeodontinae, and Leptaucheniinae, Three Subfamilies Of Oreodonts, With an Appendix to the Revision Of The Merycoidodontidae, and Part 2. Summary and Conclusions Concerning the Merycoidodontidae, Bulletin of the American Museum of Natural History 139, 1-498

Edwards P. (1976) The subfamily Leptochoerinae (Artiodactyla, Dichobunidae) of North America (Oligocene), Contributions to Geology, University of Wyoming 14 2, 99-113

Korth W. W. (1989) Aplodontid Rodents (Mammalia) from the Oligocene (Orellan and Whitneyan) Brule Formation, Nebraska, Journal of Vertebrate Paleontology 9 4, 400-414

Wang X. (1994) Phylogenetic systematics of the Hesperocyoninae (Carnivora: Canidae), Bulletin of the American Museum of Natural History 221, 1-207

Tedford R. H., et al. (1996)

Prothero D. R. (1996) Camelidae, The Terrestrial Eocene-Oligocene Transition in North America, 609-651

Prothero D. R. (1996) Hyracodontidae, The Terrestrial Eocene-Oligocene Transition in North America, 652-663

Alroy J. (2002) Synonymies and reidentifications of North American fossil mammals

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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