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Cercopidae

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Description

Cercopidae are the largest family of Cercopoidea, a xylem-feeding insect group, commonly called froghoppers or spittlebugs. They belong to the hemipteran suborder Auchenorrhyncha.

From Wikipedia article at https://en.wikipedia.org/wiki/Cercopidae, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	3763	https://www.gbif.org/species/3763
PaleoBioDB ID (PBDB)	190116	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=190116

Rank

family

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Hemiptera : Cercopidae

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Clareocercaria	Prokop et al. 2017
-	Acercaria
superorder	Pancondylognatha	Yoshizawa and Leinhard 2016
-	Condylognatha	Börner 1904
order	Hemiptera	Linnaeus 1758
suborder	Auchenorrhyncha
infraorder	Cicadomorpha	Evans 1946
superfamily	Cercopoidea	Leach 1815
family	Cercopidae	Leach 1815

Common Name

froghopper

Scientific Name

Cercopidae

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Cercopidae	family	belongs to Homoptera	stated without evidence	Tillyard, 1918
Cercopidae	family	belongs to Homoptera	stated with evidence	Zeuner, 1941
Cercopidae	family	belongs to Fulgoroidea	stated with evidence	Statz, 1950
Cercopidae	family	belongs to Cercopoidea	stated with evidence	Becker-Migdisova, 1964

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

5 (7.0%)

First Recorded Appearance

145 - 140 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

R. J. Tillyard. 1918. Permian and Triassic insects from New South Wales, in the collection of Mr. John Mitchell. The Proceedings of the Linnean Society of New South Wales 42:720-756

Common Name(s)

Froghoppers, Spittlebugs, Schuimcicaden, Froghopper

Wikipedia

https://en.wikipedia.org/wiki/Cercopidae

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Cercopinae subfamily	listed (PBDB)		25	56.0 Ma Paleocene	0 Ma Extant
Abidama genus	accepted (GBIF)	No associated record in PBDB
Aeneolamia genus	accepted (GBIF)	No associated record in PBDB
Allocercopis ✝ genus	accepted (GBIF) listed (PBDB)		2	66.0 Ma Late/Upper Cretaceous	66.0 Ma Late/Upper Cretaceous
Alluaudensia genus	accepted (GBIF)	No associated record in PBDB
Amberana genus	accepted (GBIF)	No associated record in PBDB
Ambonga genus	accepted (GBIF)	No associated record in PBDB
Anoplosnastus genus	accepted (GBIF)	No associated record in PBDB
Anyllis genus	accepted (GBIF)	No associated record in PBDB
Aracamunia genus	accepted (GBIF)	No associated record in PBDB
Asiacercopis genus	accepted (GBIF)	No associated record in PBDB
Aufidus genus	accepted (GBIF)	No associated record in PBDB
Augustohahnia genus	accepted (GBIF)	No associated record in PBDB
Baetkia genus	accepted (GBIF)	No associated record in PBDB
Baibarana genus	accepted (GBIF)	No associated record in PBDB
Bandusia genus	accepted (GBIF)	No associated record in PBDB
Blotea genus	accepted (GBIF)	No associated record in PBDB
Bourgoinrana genus	accepted (GBIF)	No associated record in PBDB
Bradypteroscarta genus	accepted (GBIF)	No associated record in PBDB
Callitettix genus	accepted (GBIF)	No associated record in PBDB
Caloscarta genus	accepted (GBIF)	No associated record in PBDB
Carachata genus	accepted (GBIF)	No associated record in PBDB
Carpentiera genus	accepted (GBIF)	No associated record in PBDB
Catrimania genus	accepted (GBIF)	No associated record in PBDB
Cercopicesa genus	accepted (GBIF)	No associated record in PBDB
Cercopis genus	accepted (GBIF) listed (PBDB)		22	56.0 Ma Paleocene	0 Ma Extant
Cercopites ✝ genus	accepted (GBIF) listed (PBDB)		3	56.0 Ma Paleocene	50.3 Ma Eocene
Chinana genus	accepted (GBIF)	No associated record in PBDB
Choconta genus	accepted (GBIF)	No associated record in PBDB
Cicadellites ✝ genus	accepted (GBIF) listed (PBDB)		5	12.7 Ma Miocene	3.60 Ma Pliocene
Clypeocarta genus	accepted (GBIF)	No associated record in PBDB
Colsa genus	accepted (GBIF)	No associated record in PBDB
Considia genus	accepted (GBIF)	No associated record in PBDB
Cosmoscarta genus	accepted (GBIF)	No associated record in PBDB
Dawsonites ✝ genus	accepted (GBIF) listed (PBDB)		1	56.0 Ma Paleocene	56.0 Ma Paleocene
Deois genus	accepted (GBIF)	No associated record in PBDB
Deoisella genus	accepted (GBIF)	No associated record in PBDB
Diabaena genus	accepted (GBIF)	No associated record in PBDB
Dulitana genus	accepted (GBIF)	No associated record in PBDB
Ectemnocarta genus	accepted (GBIF)	No associated record in PBDB
Ectemnonotops genus	accepted (GBIF)	No associated record in PBDB
Ectemnonotum genus	accepted (GBIF)	No associated record in PBDB
Eocercopidium ✝ genus	accepted (GBIF) listed (PBDB)		1	59.2 Ma Paleocene	59.2 Ma Paleocene
Eoscarta genus	accepted (GBIF)	No associated record in PBDB
Eubakeriella genus	accepted (GBIF)	No associated record in PBDB
Euglobiceps genus	accepted (GBIF)	No associated record in PBDB
Euryaulax genus	accepted (GBIF)	No associated record in PBDB
Euryliterna genus	accepted (GBIF)	No associated record in PBDB
Ferorhinella genus	accepted (GBIF)	No associated record in PBDB
Gervasiella genus	accepted (GBIF)	No associated record in PBDB
Gynopygocarta genus	accepted (GBIF)	No associated record in PBDB
Gynopygoplax genus	accepted (GBIF)	No associated record in PBDB
Haematoloma genus	accepted (GBIF)	No associated record in PBDB
Haematoscarta genus	accepted (GBIF)	No associated record in PBDB
Helioscarta genus	accepted (GBIF)	No associated record in PBDB
Hemiaufidus genus	accepted (GBIF)	No associated record in PBDB
Hemibandusia genus	accepted (GBIF)	No associated record in PBDB
Hemicercopis genus	accepted (GBIF)	No associated record in PBDB
Hemieoscarta genus	accepted (GBIF)	No associated record in PBDB
Hemiliterna genus	accepted (GBIF)	No associated record in PBDB
Hemiplagiophleboptena genus	accepted (GBIF)	No associated record in PBDB
Hemitomaspis genus	accepted (GBIF)	No associated record in PBDB
Hemitrichoscarta genus	accepted (GBIF)	No associated record in PBDB
Hemitriecphora genus	accepted (GBIF) listed (PBDB)		1	0.01 Ma Pleistocene	0 Ma Extant
Heteroliterna genus	accepted (GBIF)	No associated record in PBDB
Homalogrypota genus	accepted (GBIF)	No associated record in PBDB
Homalostethus genus	accepted (GBIF)	No associated record in PBDB
Huaina genus	accepted (GBIF)	No associated record in PBDB
Hyalotomaspis genus	accepted (GBIF)	No associated record in PBDB
Hyboscarta genus	accepted (GBIF)	No associated record in PBDB
Iphirhina genus	accepted (GBIF)	No associated record in PBDB
Irridiculum genus	accepted (GBIF)	No associated record in PBDB
Ischnorhina genus	accepted (GBIF)	No associated record in PBDB
Isozulia genus	accepted (GBIF)	No associated record in PBDB
Jacobsoniella genus	accepted (GBIF)	No associated record in PBDB
Jeanneliensia genus	accepted (GBIF)	No associated record in PBDB
Kanaima genus	accepted (GBIF)	No associated record in PBDB
Kanoscarta genus	accepted (GBIF)	No associated record in PBDB
Kanozata genus	accepted (GBIF)	No associated record in PBDB
Korobona genus	accepted (GBIF)	No associated record in PBDB
Kotozata genus	accepted (GBIF)	No associated record in PBDB
Laccogrypota genus	accepted (GBIF)	No associated record in PBDB
Lamprochlamys genus	accepted (GBIF)	No associated record in PBDB
Lehina genus	accepted (GBIF)	No associated record in PBDB
Leptataspis genus	accepted (GBIF)	No associated record in PBDB
Leptoliterna genus	accepted (GBIF)	No associated record in PBDB
Leptynis genus	accepted (GBIF)	No associated record in PBDB
Lieftinckana genus	accepted (GBIF)	No associated record in PBDB
Liorhinella genus	accepted (GBIF)	No associated record in PBDB
Literna genus	accepted (GBIF)	No associated record in PBDB
Lithecphora ✝ genus	accepted (GBIF) listed (PBDB)		4	37.2 Ma Eocene	37.2 Ma Eocene
Locris genus	accepted (GBIF)	No associated record in PBDB
Locrites ✝ genus	accepted (GBIF) listed (PBDB)		5	37.2 Ma Eocene	12.7 Ma Miocene
Luederwaldtia genus	accepted (GBIF)	No associated record in PBDB
Lujana genus	accepted (GBIF)	No associated record in PBDB
Machadoa genus	accepted (GBIF)	No associated record in PBDB
Mahanarva genus	accepted (GBIF)	No associated record in PBDB
Makonaima genus	accepted (GBIF)	No associated record in PBDB
Marcion genus	accepted (GBIF)	No associated record in PBDB
Maxantonia genus	accepted (GBIF)	No associated record in PBDB
Megacercopis ✝ genus	accepted (GBIF) listed (PBDB)		1	20.4 Ma Miocene	20.4 Ma Miocene
Megastethodon genus	accepted (GBIF)	No associated record in PBDB
Microclimax genus	accepted (GBIF)	No associated record in PBDB
Microlaqueus genus	accepted (GBIF)	No associated record in PBDB
Microsargane genus	accepted (GBIF)	No associated record in PBDB
Mioscarta genus	accepted (GBIF)	No associated record in PBDB
Monecphora genus	accepted (GBIF)	No associated record in PBDB
Moultoniella genus	accepted (GBIF)	No associated record in PBDB
Neaenus genus	accepted (GBIF)	No associated record in PBDB
Neocercopis genus	accepted (GBIF)	No associated record in PBDB
Neolaccogrypota genus	accepted (GBIF)	No associated record in PBDB
Neomonecphora genus	accepted (GBIF)	No associated record in PBDB
Neoporpacella genus	accepted (GBIF)	No associated record in PBDB
Neosphenorhina genus	accepted (GBIF)	No associated record in PBDB
Nesaphrogeneia genus	accepted (GBIF)	No associated record in PBDB
Nisocercopis ✝ genus	accepted (GBIF) listed (PBDB)		1	66.0 Ma Late/Upper Cretaceous	66.0 Ma Late/Upper Cretaceous
Notozulia genus	accepted (GBIF)	No associated record in PBDB
Ocoaxo genus	accepted (GBIF)	No associated record in PBDB
Okiscarta genus	accepted (GBIF)	No associated record in PBDB
Olcotomaspis genus	accepted (GBIF)	No associated record in PBDB
Opistharsostethus genus	accepted (GBIF)	No associated record in PBDB
Orodamnis genus	accepted (GBIF)	No associated record in PBDB
Orthorhinella genus	accepted (GBIF)	No associated record in PBDB
Oxymegaspis genus	accepted (GBIF)	No associated record in PBDB
Oxypeltarion genus	accepted (GBIF)	No associated record in PBDB
Pachacanthocnemis genus	accepted (GBIF)	No associated record in PBDB
Palecphora ✝ genus	accepted (GBIF) listed (PBDB)		13	56.0 Ma Paleocene	37.2 Ma Eocene
Panabrus genus	accepted (GBIF)	No associated record in PBDB
Paphnutius genus	accepted (GBIF)	No associated record in PBDB
Paracercopis genus	accepted (GBIF)	No associated record in PBDB
Parafitopteryx ✝ genus	accepted (GBIF) listed (PBDB)		1	28.1 Ma Oligocene	28.1 Ma Oligocene
Paraliterna genus	accepted (GBIF)	No associated record in PBDB
Paralocris genus	accepted (GBIF)	No associated record in PBDB
Paramioscarta genus	accepted (GBIF)	No associated record in PBDB
Paramonecphora genus	accepted (GBIF)	No associated record in PBDB
Parapisidice genus	accepted (GBIF)	No associated record in PBDB
Paratrichoscarta genus	accepted (GBIF)	No associated record in PBDB
Petyllis genus	accepted (GBIF)	No associated record in PBDB
Phlebarcys genus	accepted (GBIF)	No associated record in PBDB
Phymatostetha genus	accepted (GBIF)	No associated record in PBDB
Pisianax genus	accepted (GBIF)	No associated record in PBDB
Pisidice genus	accepted (GBIF)	No associated record in PBDB
Plagiophleboptena genus	accepted (GBIF)	No associated record in PBDB
Platynotoscarta genus	accepted (GBIF)	No associated record in PBDB
Poeciloterpa genus	accepted (GBIF)	No associated record in PBDB
Pogonorhinella genus	accepted (GBIF)	No associated record in PBDB
Porpacella genus	accepted (GBIF)	No associated record in PBDB
Prinecphora ✝ genus	accepted (GBIF) listed (PBDB)		2	37.2 Ma Eocene	37.2 Ma Eocene
Prosapia genus	accepted (GBIF)	No associated record in PBDB
Pseudaufidus genus	accepted (GBIF)	No associated record in PBDB
Pseudeoscarta genus	accepted (GBIF)	No associated record in PBDB
Pseudocercopis genus	accepted (GBIF)	No associated record in PBDB
Pseudomachaerota genus	accepted (GBIF)	No associated record in PBDB
Ptychotrichos genus	accepted (GBIF)	No associated record in PBDB
Ptyelus genus	doubtful (GBIF) listed (PBDB)		5	33.9 Ma Oligocene	0 Ma Extant
Radioscarta genus	accepted (GBIF)	No associated record in PBDB
Rhinastria genus	accepted (GBIF)	No associated record in PBDB
Rhinaulax genus	accepted (GBIF)	No associated record in PBDB
Schistogonia genus	accepted (GBIF)	No associated record in PBDB
Serapita genus	accepted (GBIF)	No associated record in PBDB
Simeliria genus	accepted (GBIF)	No associated record in PBDB
Simorhina genus	accepted (GBIF)	No associated record in PBDB
Sinocercopis ✝ genus	accepted (GBIF) listed (PBDB)		2	133 Ma Early/Lower Cretaceous	125 Ma Early/Lower Cretaceous
Sinopia genus	accepted (GBIF)	No associated record in PBDB
Sphenoclypeana genus	accepted (GBIF)	No associated record in PBDB
Sphenorhina genus	accepted (GBIF)	No associated record in PBDB
Stenecphora ✝ genus	accepted (GBIF) listed (PBDB)		1	56.0 Ma Paleocene	56.0 Ma Paleocene
Stenolocris ✝ genus	accepted (GBIF) listed (PBDB)		1	56.0 Ma Paleocene	56.0 Ma Paleocene
Straelenia genus	accepted (GBIF)	No associated record in PBDB
Suracarta genus	accepted (GBIF)	No associated record in PBDB
Tadascarta genus	accepted (GBIF)	No associated record in PBDB
Tapaiuna genus	accepted (GBIF)	No associated record in PBDB
Telogmometopius genus	accepted (GBIF)	No associated record in PBDB
Thoodzata genus	accepted (GBIF)	No associated record in PBDB
Tiodus genus	accepted (GBIF)	No associated record in PBDB
Tomaspis genus	accepted (GBIF)	No associated record in PBDB
Tomaspisina genus	accepted (GBIF)	No associated record in PBDB
Tomaspisinella genus	accepted (GBIF)	No associated record in PBDB
Trichoscarta genus	accepted (GBIF)	No associated record in PBDB
Triecphorella genus	accepted (GBIF)	No associated record in PBDB
Tropidorhinella genus	accepted (GBIF)	No associated record in PBDB
Tunaima genus	accepted (GBIF)	No associated record in PBDB
Typeschata genus	accepted (GBIF)	No associated record in PBDB
Urubaxia genus	accepted (GBIF)	No associated record in PBDB
Vigilantius genus	accepted (GBIF)	No associated record in PBDB
Villiersana genus	accepted (GBIF)	No associated record in PBDB
Vorago genus	accepted (GBIF)	No associated record in PBDB
Zuata genus	accepted (GBIF)	No associated record in PBDB
Zulia genus	accepted (GBIF)	No associated record in PBDB
Cosmocarta dimidrata species	accepted (GBIF)	No associated record in PBDB
Wawi mehi species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Meloe hoernesi Giebel, 1856	GBIF	species	Animalia : Arthropoda : Insecta : Hemiptera : Cercopidae : *Meloe hoernesi*

Synonymy List

Year	Name and Author
1815	Cercopidae Leach
1856	Meloe hoernesi Giebel p. 106
1918	Cercopidae Tillyard
1921	Cercopidae Cockerell
1941	Cercopidae Zeuner p. 88
1950	Cercopidae Statz p. 5

Year	Name and Author
1964	Cercopidae Becker-Migdisova p. 7
1992	Cercopidae Hamilton p. 428
2012	Cercopidae Cryan and Urban
2018	Cercopidae Szwedo p. 110
2019	Cercopidae Szwedo p. 375

References

Tillyard R. J. (1918) Permian and Triassic insects from New South Wales, in the collection of Mr. John Mitchell, The Proceedings of the Linnean Society of New South Wales 42, 720-756

Giebel C. G. Die Insecten und Spinnen der Vorwelt mit steter Berücksichtigung der lebenden Insekten und Spinnen, Die Fauna der Vorwelt 2, 1-511

Cockerell T. D. A. (1921) Eocene Insects from the Rocky Mountains, Proceedings of the United States National Museum 57, 233-260

Zeuner F. E. (1941) The Eocene Insects of the Ardtun Beds, Isle of Mull, Scotland, The Annals and Magazine of Natural History, Eleventh Series 7, 82-100

Statz G. (1950) Cicadariae (Zikaden) aus den Oberoligocanen Ablagerungen von Rott, Palaeontographica Abteilung A 98, 1-46

Becker-Migdisova E. E. (1964) Treticnye ravokrylye Stravropol'ja, Trudy Paleontologicheskogo Instituta Moskva 54, 1-104

Hamilton K. G. A. (1992) Lower Cretaceous Homoptera from the Koonwarra Fossil Bed in Australia, with a New Superfamily and Synopsis of Mesozoic Homoptera, Annals of the Entomological Society of America 85 4, 423-430

Szwedo J. (2018) The unity, diversity and conformity of bugs (Hemiptera) through time, Earth and Environmental Science Transactions of the Royal Society of Edinburgh 107, 109-128

Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera

Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera

Cryan, Jason R., and Gavin J. Svenson, 2010: Family-level relationshipts of the spittlebugs and froghoppers (Hemiptera: Cicadomorpha: Cercopoidea). Systematic Entomology, vol. 35, no. 3. 393-415. - via Integrated Taxonomic Information System (ITIS)

McGavin, George C., 1993: null. Bugs of the World. 192. - via Integrated Taxonomic Information System (ITIS)

Krepinsky, O. (1978). Saltmarsh insect survey of 1978. In G. W. Gratto (Ed.) Some aspects of the flora and fauna of the western Minas Basin and Minas Channel. (pp. 10-28). Acadia University. Wolfville, Nova Scotia. - via World Register of Marine Species

Hamilton, K.G. Andrew (2001) A new family of froghoppers from the American tropics (Hemiptera: Cercopoidea: Epipygidae). Biodiversity 2 (3): 15-21. - via Dyntaxa. Svensk taxonomisk databas

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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