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Clusiidae

Description

Clusiidae or "druid flies" is a family of small (~ 3.5 mm), thin, yellow to black acalyptrate flies with a characteristic antenna (The second segment of the antennae has a triangular projection over the third segment when viewed from the outside) and with the wing usually partially infuscated. They have a cylindrical body. The head is round, the vertical plate reaches the anterior margin of the frons and the vibrissae on the head are large. The costa is interrupted near subcosta and the latter developed throughout length. Larvae are found in the bark of trees, the flies on trunks.The larvae are notable for their ability to jump. Males of many species in the subfamily Clusiodinae have been observed while engaged in lekking behaviour. There are hundreds of species in 14 genera found in all the Ecoregions, although most species occur in tropical regions. The type genus is Clusia Haliday, 1838.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)7290https://www.gbif.org/species/7290
PaleoBioDB ID (PBDB)139325https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=139325
Rankfamily
Taxonomy (GBIF,PBDB)Life : Animalia : Arthropoda : Insecta : Diptera : Clusiidae
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-Bilateria
-EubilateriaAx 1987
-ProtostomiaGrobben 1908
-Ecdysozoa
-Panarthropoda
phylumArthropodaLatreille 1829
subphylumMandibulata
-Pancrustacea
superclassHexapodaLatreille 1825
-CercofilataKukalová-Peck 1987
classInsectaLinnaeus 1758
-Dicondylia
-ParanotaliaSroka et al. 2014
subclassPterygotaBrauer 1885
-NeopterygotaCrampton 1924
infraclassNeopteraMartynov 1923
-Holometabola
-Aparaglossata
-Mecopterida
superorderAntliophora
orderDipteraLinnaeus 1758
-NeodipteraMichelsen 1994
suborderBrachyceraZetterstedt 1842
-Heterodactyla
-EremoneuraGrimaldi and Cumming 1999
suborderCyclorrhapha
-SchizophoraBecher 1882
familyClusiidaeFrey 1921
Common Namedruid fly
Scientific NameClusiidae
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Clusiidaefamilybelongs to Brachycerastated without evidenceCarpenter, 1992
Clusiidaefamilybelongs to Schizophorastated with evidenceWiegmann et al., 2011
Status (PBDB)extant
Taxon Size (PBDB)10
Extant Size (PBDB)4 (40%)
First Recorded Appearance38.0 - 33.9 Ma
Paleogene
Environmentterrestrial (based on Insecta)
Motilityactively mobile (based on Arthropoda)
Taphonomychitin (based on Insecta)
Primary Reference (PBDB)R. Frey. 1921. Studien uber den Bau des Mundes der niederen Diptera Schizophora nebst Bemerkungen uber die Systematik dieser Dipterengruppe. Acta Societatis pro Fauna et Flora Fennica 48:1-245
Common Name(s) Druid Fly, Clusiid Flies, Träflugor
Wikipediahttps://en.wikipedia.org/wiki/Clusiidae

External Images

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Clusiinae
subfamily
listed (PBDB)0 Ma
Extant
Clusiodinae
subfamily
listed (PBDB)638.0 Ma
Eocene
0 Ma
Extant
Alloclusia
genus
accepted (GBIF)No associated record in PBDB
Allometopon
genus
accepted (GBIF)No associated record in PBDB
Amuroclusia
genus
accepted (GBIF)No associated record in PBDB
Apiochaeta
genus
accepted (GBIF)No associated record in PBDB
Chaetoclusia
genus
accepted (GBIF)No associated record in PBDB
Chaetoclusiella
genus
accepted (GBIF)No associated record in PBDB
Clusia
genus
accepted (GBIF)No associated record in PBDB
Clusiodes
genus
accepted (GBIF)
listed (PBDB)
128.1 Ma
Oligocene
0 Ma
Extant
Craspedochaeta
genus
accepted (GBIF)No associated record in PBDB
Czernyola
genus
accepted (GBIF)No associated record in PBDB
Electroclusiodes
genus
accepted (GBIF)
listed (PBDB)
538.0 Ma
Eocene
38.0 Ma
Eocene
Hendelia
genus
accepted (GBIF)No associated record in PBDB
Heteromeringia
genus
accepted (GBIF)No associated record in PBDB
Hteromeringia
genus
accepted (GBIF)No associated record in PBDB
Melanoclusia
genus
accepted (GBIF)No associated record in PBDB
Microclusiaria
genus
accepted (GBIF)No associated record in PBDB
Phylloclusia
genus
accepted (GBIF)No associated record in PBDB
Procerosoma
genus
accepted (GBIF)No associated record in PBDB
Prohendelia
genus
accepted (GBIF)No associated record in PBDB
Sobarocephala
genus
accepted (GBIF)No associated record in PBDB
Sobarocephaloides
genus
accepted (GBIF)No associated record in PBDB
Tetrameringia
genus
accepted (GBIF)No associated record in PBDB
Tranomeringia
genus
accepted (GBIF)No associated record in PBDB
Trichoclusia
genus
accepted (GBIF)No associated record in PBDB
Xenanthomyza
genus
accepted (GBIF)
listed (PBDB)
138.0 Ma
Eocene
38.0 Ma
Eocene
Chaetochlusia nigromaculata
species
accepted (GBIF)No associated record in PBDB
Chaetochlusia quadrivittata
species
accepted (GBIF)No associated record in PBDB
Chaetolusia bakeri
species
accepted (GBIF)No associated record in PBDB
Cludiodes nitida
species
accepted (GBIF)No associated record in PBDB

Synonymy List

YearName and Author
1921Clusiidae Frey p. 29
1992Clusiidae Carpenter
2011Clusiidae Wiegmann et al.

References

Carpenter F. M. (1992) , Treatise on Invertebrate Paleontology Part R, Arthropoda 4: Superclass Hexapoda 3/4, 1-655
Wiegmann B. M., Trautwein M. D., et al (2011) Episodic radiations in the fly tree of life, Proceedings of the National Academy of Sciences 108, 5690-5695
Gärdenfors, Ulf, Hall, Ragnar, Hallingbäck, Tomas, Hansson, Hans G. & Hedström, Lars (2003) Djur, svampar och växter i Sverige 2003. Förteckning över antal arter per familj. ArtDatabanken Rapporterar 5, ArtDatabanken, Uppsala. (reviderad excelfil med svenska djurfamiljer). - via Dyntaxa. Svensk taxonomisk databas
Thompson, F.C., Evenhuis, N.L. & Sabrosky, C.W. (1999) Family-Group Names in Diptera. An annotated catalog. Myia 10: 1-360 - via Dyntaxa. Svensk taxonomisk databas
Systema Dipterorum (2013) http://www.diptera.org/About.php - via Dyntaxa. Svensk taxonomisk databas
Teskey, H. J.,McAlpine, J. F.,Soós, Á,Shewell, G. E.,Wood, D. M.,Vockeroth, J. R.,Peterson, B. V. (1987.03.31) Chapter 70. Clusiidae - via The Diptera Site
Benton, M.J. (ed). (1993). The Fossil Record 2. Chapman & Hall, London, 845 pp. - via The Interim Register of Marine and Nonmarine Genera
Parker, S.P. (ed). (1982). Synopsis and Classification of Living Organisms. McGraw-Hill, New York. 2 volumes. - via The Interim Register of Marine and Nonmarine Genera
Papp, L.,Sasakawa, M.,Darvas, B. (1998) 3.16. Family Clusiidae - via The Diptera Site
Thompson, F.C., Evenhuis, N.L. & Sabrosky, C.W. (1999) Family-Group Names in Diptera. An annotated catalog. Myia 10: 1-360 - via Dyntaxa. Svensk taxonomisk databas
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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