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Vancleavea

Description

Vancleavea is a genus of extinct, armoured, non-archosaurian archosauriforms from the Late Triassic of western North America. The type and only known species is V. campi, named by Robert Long & Phillip A Murry in 1995. At that time, the genus was only known from fragmentary bones including osteoderms and vertebrae. However, since then many more fossils have been found, including a pair of nearly complete skeletons discovered in 2002. These finds have shown that members of the genus were bizarre semiaquatic reptiles. Vancleavea individuals had short snouts with large, fang-like teeth, and long bodies with small limbs. They were completely covered with bony plates known as osteoderms, which came in several different varieties distributed around the body. Phylogenetic analyses by professional paleontologists have shown that Vancleavea was an archosauriform, part of the lineage of reptiles that would lead to archosaurs such as dinosaurs and crocodilians. Vancleavea lacks certain traits which are present in most other archosauriforms, most notably the antorbital, mandibular and supratemporal fenestrae, which are weight-saving holes in the skulls of other taxa. However, other features clearly support its archosauriform identity, including a lack of intercentra, the presence of osteoderms, an ossified laterosphenoid, and several adaptations of the femur and ankle bones. In 2016, a new genus of archosauriform, Litorosuchus, was described. This genus resembled both Vancleavea and more typical archosauriforms in different respects, allowing Litorosuchus to act as a transitional fossil linking Vancleavea to less aberrant archosauriforms.


Source Data
SourceIDLink
Global Biodiversity Information Facility ID (GBIF)4822422https://www.gbif.org/species/4822422
PaleoBioDB ID (PBDB)69042https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=69042
Rankgenus
Taxonomy (GBIF)Life : Animalia : Chordata : Reptilia : Doswelliidae : Vancleavea
Taxonomy (PBDB)Life : Animalia : Chordata : Reptilia : Eosuchia : Doswelliidae : Vancleavea
Taxonomic Status (GBIF)accepted
Classification
(PBDB,GBIF)
RankNameAuthor
-Eukaryota
-OpisthokontaCavalier-Smith 1987
kingdomAnimalia
-TriploblasticaLankester 1877
-NephrozoaJondelius et al. 2002
-DeuterostomiaGrobben 1908
phylumChordataHaeckel 1847
subphylumVertebrataLamarck 1801
superclassGnathostomataGegenbauer 1874
-Osteichthyes
-SarcopterygiiRomer 1955
subclassDipnotetrapodomorpha
subclassTetrapodomorpha
-Tetrapoda
-ReptiliomorphaSäve-Söderbergh 1934
-AnthracosauriaSäve-Söderbergh 1934
-Batrachosauria
-Cotylosauria
-AmniotaHaeckel 1866
-SauropsidaHuxley 1864
classReptiliaLaurenti 1768
subclassEureptilia
-RomeriidaGauthier et al. 1988
-Diapsida
-Eosuchia
-NeodiapsidaBenton 1985
-SauriaGauthier 1984
-Archosauromorpha
-CrocopodaEzcurra 2016
-ArchosauriformesGauthier 1986
-EucrocopodaEzcurra 2016
-ProterochampsiaKischlat 2000
-Doswelliidae
genusVancleaveaLong & Murry 1995
Scientific NameVancleavea Long & Murry, 1995
Name Published InBull N M Mus Nat Hist Sci 4
Opinions (PBDB)
NameRankOpinionEvidenceAuthor
Vancleaveagenusbelongs to Neodiapsidastated with evidenceLong and Murry, 1995
Vancleaveagenusbelongs to ArchosauromorphaimpliedIrmis, 2005
Vancleaveagenusbelongs to Archosauriformesstated with evidenceParker and Barton, 2008
Vancleaveagenusbelongs to Doswelliidaestated with evidenceEzcurra, 2016
Status (PBDB)extinct
Taxon Size (PBDB)2
First Recorded Appearance237 - 228 Ma
Late/Upper Triassic
Last Recorded Appearance209 - 201 Ma
Mesozoic
Environmentfreshwater
Motilityactively mobile (based on Osteichthyes)
Dietcarnivore
Taphonomyphosphatic (based on Vertebrata)
Primary Reference (PBDB)R. A. Long and P. A. Murry. 1995. Late Triassic (Carnian and Norian) tetrapods from the southwestern United States. New Mexico Museum of Natural History and Science Bulletin 4:1-254
Wikipediahttps://en.wikipedia.org/wiki/Vancleavea

Fossil Distribution

Subtaxa

NameStatusCommon Name(s)Fossil OccurrencesOldestYoungest
Vancleavea campi
species
accepted (GBIF)
listed (PBDB)
19237 Ma
Middle Triassic
209 Ma
Late/Upper Triassic

Synonymy List

YearName and Author
1995Vancleavea Long and Murry
2005Vancleavea Irmis p. 71
2008Vancleavea Parker and Barton
2009Vancleavea Nesbitt et al.
2011Vancleavea Nesbitt p. 191 fig. 51
2012Vancleavea Spielmann and Lucas
2016Vancleavea Ezcurra

References

Long R. A., Murry P. A. (1995) Late Triassic (Carnian and Norian) tetrapods from the southwestern United States, New Mexico Museum of Natural History and Science Bulletin 4, 1-254
Irmis R. B. (2005) The vertebrate fauna of the Upper Triassic Chinle Formation in northern Arizona, In S. J. Nesbitt, W. G. Parker, & R. B. Irmis (eds.), Guidebook to the Triassic Formations of the Colorado Plateau in Northern Arizona: Geology, Paleontology, and History. Mesa Southwest Museum Bulletin 9, 63-88
Parker W. G., Barton B. J. (2008) New information on the Upper Triassic archosauriform Vancleavea campi based on new material from the Chinle Formation of Arizona, Palaeontologia Electronica 11 3, 14A
Nesbitt S. J., Stocker M. R., et al (2009) The osteology and relationships of Vancleavea campi (Reptilia: Archosauriformes), Zoological Journal of the Linnean Society 157, 814-864 doi:10.1111/j.1096-3642.2009.00530.x
Nesbitt S. J. (2011) The early evolution of archosaurs: relationships and the origin of major clades, Bulletin of the American Museum of Natural History 353, 1-292
Spielmann J. A., Lucas S. G. (2012) Tetrapod fauna of the Upper Triassic Redonda formation, East-Central New Mexico: the characteristic assemblage of the Apachean land-vertebrate faunachron, New Mexico Museum of Natural History and Science, Bulletin 55, 1-119
Ezcurra M. D. (2016) The phylogenetic relationships of basal archosauromorphs, with an emphasis on the systematics of proterosuchian archosauriforms, PeerJ 4, e1778
Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera
Bull N M Mus Nat Hist Sci 4 - via The Interim Register of Marine and Nonmarine Genera
Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!
 
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