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Cassida

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Description

Cassida is a large Old World genus of tortoise beetles in the subfamily Cassidinae. Several species of Cassida are important agricultural pests, in particular C. vittata and C. nebulosa on sugar beet and spinach. The thistle tortoise beetle (Cassida rubiginosa) has been used as a biological control agent against Canada thistle.

From Wikipedia article at https://en.wikipedia.org/wiki/Cassida, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	1048148	https://www.gbif.org/species/1048148
PaleoBioDB ID (PBDB)	207282	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=207282

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : Cassida

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Polyphaga	Emery 1886
infraorder	Cucujiformia	Lameere 1938
-	Phytophaga
superfamily	Chrysomeloidea	Latreille 1802
family	Chrysomelidae	Latreille 1802
subfamily	Cassidinae	Gyllenhal 1813
tribe	Cassidini	Gyllenhal 1813
genus	Cassida	Linnæus 1758

Scientific Name

Cassida Linnaeus, 1758

Name Published In

Linnæus, Carolus. 1758. Systema naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Laurentii Salvii, Holmiae. Vol. Tomus I, Editio decima, reformata: i-ii, 1-824.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Cassida	genus	belongs to Coleoptera	stated with evidence	Linnaeus, 1758
Cassida	genus	belongs to Eleuterata	stated with evidence	Fabricius, 1775
Cassida	genus	belongs to Coleoptera	stated with evidence	Müller, 1776
Cassida	genus	belongs to Eleuterata	stated with evidence	Fabricius, 1798
Cassida	genus	belongs to Galerucae	stated with evidence	Latreille, 1802
Cassida	genus	belongs to Cassidini	stated with evidence	Bouchard et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

1 (14%)

First Recorded Appearance

56.0 - 47.8 Ma
Eocene

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

C. Linnaeus. 1758. Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1:1-824

Common Name(s)

Schildpadtorren, Schildtorretjes, Schildpadtorretjes, Tortoise Beetles

Wikipedia

https://en.wikipedia.org/wiki/Cassida

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Cassida abamita species	accepted (GBIF)	No associated record in PBDB
Cassida achardi species	accepted (GBIF)	No associated record in PBDB
Cassida acutangula species	accepted (GBIF)	No associated record in PBDB
Cassida aethiopica species	accepted (GBIF)	No associated record in PBDB
Cassida agilis species	accepted (GBIF)	No associated record in PBDB
Cassida albertisi species	accepted (GBIF)	No associated record in PBDB
Cassida algirica species	accepted (GBIF)	No associated record in PBDB
Cassida alpina species	accepted (GBIF)	No associated record in PBDB
Cassida alticola species	accepted (GBIF)	No associated record in PBDB
Cassida altiuscula species	accepted (GBIF)	No associated record in PBDB
Cassida amaranthica species	accepted (GBIF)	No associated record in PBDB
Cassida ambrica species	accepted (GBIF)	No associated record in PBDB
Cassida amorifica species	accepted (GBIF)	No associated record in PBDB
Cassida amurensis species	accepted (GBIF)	No associated record in PBDB
Cassida ancorifera species	accepted (GBIF)	No associated record in PBDB
Cassida andapaensis species	accepted (GBIF)	No associated record in PBDB
Cassida andreinii species	accepted (GBIF)	No associated record in PBDB
Cassida andrewesi species	accepted (GBIF)	No associated record in PBDB
Cassida angusta species	accepted (GBIF)	No associated record in PBDB
Cassida angustifrons species	accepted (GBIF)	No associated record in PBDB
Cassida antoni species	accepted (GBIF)	No associated record in PBDB
Cassida appluda species	accepted (GBIF)	No associated record in PBDB
Cassida astrolabiana species	accepted (GBIF)	No associated record in PBDB
Cassida atrata species	accepted (GBIF)	No associated record in PBDB
Cassida atripes species	accepted (GBIF)	No associated record in PBDB
Cassida atrofemorata species	accepted (GBIF)	No associated record in PBDB
Cassida atrorubra species	accepted (GBIF)	No associated record in PBDB
Cassida atrosignata species	accepted (GBIF)	No associated record in PBDB
Cassida aureola species	accepted (GBIF)	No associated record in PBDB
Cassida auropustulata species	accepted (GBIF)	No associated record in PBDB
Cassida aurora species	accepted (GBIF)	No associated record in PBDB
Cassida australica species	accepted (GBIF)	No associated record in PBDB
Cassida avia species	accepted (GBIF)	No associated record in PBDB
Cassida azurea species	accepted (GBIF)	No associated record in PBDB
Cassida bakeri species	accepted (GBIF)	No associated record in PBDB
Cassida basicollis species	accepted (GBIF)	No associated record in PBDB
Cassida basilana species	accepted (GBIF)	No associated record in PBDB
Cassida bella species	accepted (GBIF)	No associated record in PBDB
Cassida belli species	accepted (GBIF)	No associated record in PBDB
Cassida belliformis species	accepted (GBIF)	No associated record in PBDB
Cassida benguelica species	accepted (GBIF)	No associated record in PBDB
Cassida beniowskii species	accepted (GBIF)	No associated record in PBDB
Cassida bergeali species	accepted (GBIF)	No associated record in PBDB
Cassida berolinensis species	accepted (GBIF)	No associated record in PBDB
Cassida bezdeki species	accepted (GBIF)	No associated record in PBDB
Cassida bicallosa species	accepted (GBIF)	No associated record in PBDB
Cassida biguttata species	accepted (GBIF)	No associated record in PBDB
Cassida binorbis species	accepted (GBIF)	No associated record in PBDB
Cassida blancheti ✝ species	accepted (GBIF) listed (PBDB)		5	28.1 Ma Oligocene	12.7 Ma Miocene
Cassida blanda species	accepted (GBIF)	No associated record in PBDB
Cassida brevis species	accepted (GBIF)	No associated record in PBDB
Cassida brooksi species	accepted (GBIF)	No associated record in PBDB
Cassida brunnea species	accepted (GBIF)	No associated record in PBDB
Cassida butterwecki species	accepted (GBIF)	No associated record in PBDB
Cassida callosicollis species	accepted (GBIF)	No associated record in PBDB
Cassida calvaria species	accepted (GBIF)	No associated record in PBDB
Cassida camerunensis species	accepted (GBIF)	No associated record in PBDB
Cassida canaliculata species	accepted (GBIF)	No associated record in PBDB
Cassida capensis species	accepted (GBIF)	No associated record in PBDB
Cassida catenata species	accepted (GBIF)	No associated record in PBDB
Cassida ceylonica species	accepted (GBIF)	No associated record in PBDB
Cassida chapuisi species	accepted (GBIF)	No associated record in PBDB
Cassida cherenica species	accepted (GBIF)	No associated record in PBDB
Cassida cherrapunjiensis species	accepted (GBIF)	No associated record in PBDB
Cassida chiangmaiensis species	accepted (GBIF)	No associated record in PBDB
Cassida chrysanthemoides species	accepted (GBIF)	No associated record in PBDB
Cassida circassica species	accepted (GBIF)	No associated record in PBDB
Cassida circumdata species	accepted (GBIF)	No associated record in PBDB
Cassida circumflexa species	accepted (GBIF)	No associated record in PBDB
Cassida circumsepta species	accepted (GBIF)	No associated record in PBDB
Cassida clavosecui species	accepted (GBIF)	No associated record in PBDB
Cassida coagulata species	accepted (GBIF)	No associated record in PBDB
Cassida coelebs species	accepted (GBIF)	No associated record in PBDB
Cassida cognobilis species	accepted (GBIF)	No associated record in PBDB
Cassida collucens species	accepted (GBIF)	No associated record in PBDB
Cassida compuncta species	accepted (GBIF)	No associated record in PBDB
Cassida concallescens species	accepted (GBIF)	No associated record in PBDB
Cassida concha species	accepted (GBIF)	No associated record in PBDB
Cassida conchyliata species	accepted (GBIF)	No associated record in PBDB
Cassida consobrina species	accepted (GBIF)	No associated record in PBDB
Cassida conspurcata species	accepted (GBIF)	No associated record in PBDB
Cassida corallina species	accepted (GBIF)	No associated record in PBDB
Cassida corbetti species	accepted (GBIF)	No associated record in PBDB
Cassida cordula species	accepted (GBIF)	No associated record in PBDB
Cassida corollata species	accepted (GBIF)	No associated record in PBDB
Cassida coronula species	accepted (GBIF)	No associated record in PBDB
Cassida corruptrix species	accepted (GBIF)	No associated record in PBDB
Cassida cristobalensis species	accepted (GBIF)	No associated record in PBDB
Cassida crucifera species	accepted (GBIF)	No associated record in PBDB
Cassida culminis species	accepted (GBIF)	No associated record in PBDB
Cassida currax species	accepted (GBIF)	No associated record in PBDB
Cassida decolorata species	accepted (GBIF)	No associated record in PBDB
Cassida deflorata species	accepted (GBIF)	No associated record in PBDB
Cassida dehradunensis species	accepted (GBIF)	No associated record in PBDB
Cassida delenifica species	accepted (GBIF)	No associated record in PBDB
Cassida delesserti species	accepted (GBIF)	No associated record in PBDB
Cassida deltoides species	accepted (GBIF)	No associated record in PBDB
Cassida dembickyi species	accepted (GBIF)	No associated record in PBDB
Cassida denticollis species	accepted (GBIF)	No associated record in PBDB
Cassida denticulata species	accepted (GBIF)	No associated record in PBDB
Cassida depicta species	accepted (GBIF)	No associated record in PBDB
Cassida derasa species	accepted (GBIF)	No associated record in PBDB
Cassida desultrix species	accepted (GBIF)	No associated record in PBDB
Cassida devalaensis species	accepted (GBIF)	No associated record in PBDB
Cassida devylderi species	accepted (GBIF)	No associated record in PBDB
Cassida diomma species	accepted (GBIF)	No associated record in PBDB
Cassida diops species	accepted (GBIF)	No associated record in PBDB
Cassida discalis species	accepted (GBIF)	No associated record in PBDB
Cassida distinguenda species	accepted (GBIF)	No associated record in PBDB
Cassida dohertyi species	accepted (GBIF)	No associated record in PBDB
Cassida dolens species	accepted (GBIF)	No associated record in PBDB
Cassida dorsalis species	accepted (GBIF)	No associated record in PBDB
Cassida dorsonotata species	accepted (GBIF)	No associated record in PBDB
Cassida dorsovittata species	accepted (GBIF)	No associated record in PBDB
Cassida drakensbergensis species	accepted (GBIF)	No associated record in PBDB
Cassida dufouri ✝ species	accepted (GBIF) listed (PBDB)		1	3.60 Ma Pliocene	3.60 Ma Pliocene
Cassida dulcis species	accepted (GBIF)	No associated record in PBDB
Cassida ellipticollis species	accepted (GBIF)	No associated record in PBDB
Cassida elongata species	accepted (GBIF)	No associated record in PBDB
Cassida eluta species	accepted (GBIF)	No associated record in PBDB
Cassida enervis species	accepted (GBIF)	No associated record in PBDB
Cassida eoa species	accepted (GBIF)	No associated record in PBDB
Cassida excaecata species	accepted (GBIF)	No associated record in PBDB
Cassida exilis species	accepted (GBIF)	No associated record in PBDB
Cassida eximia species	accepted (GBIF)	No associated record in PBDB
Cassida expansa species	accepted (GBIF)	No associated record in PBDB
Cassida expressa species	accepted (GBIF)	No associated record in PBDB
Cassida fausti species	accepted (GBIF)	No associated record in PBDB
Cassida feae species	accepted (GBIF)	No associated record in PBDB
Cassida ferranti species	accepted (GBIF)	No associated record in PBDB
Cassida ferruginea species	accepted (GBIF)	No associated record in PBDB
Cassida flaveola species	accepted (GBIF)	No associated record in PBDB
Cassida flavoguttata species	accepted (GBIF)	No associated record in PBDB
Cassida flavoscutata species	accepted (GBIF)	No associated record in PBDB
Cassida flavosignata species	accepted (GBIF)	No associated record in PBDB
Cassida foveolatipennis species	accepted (GBIF)	No associated record in PBDB
Cassida franklinmuelleri species	accepted (GBIF)	No associated record in PBDB
Cassida frontalis species	accepted (GBIF)	No associated record in PBDB
Cassida fukhanica species	accepted (GBIF)	No associated record in PBDB
Cassida fumida species	accepted (GBIF)	No associated record in PBDB
Cassida fuscomacula species	accepted (GBIF)	No associated record in PBDB
Cassida fuscopunctata species	accepted (GBIF)	No associated record in PBDB
Cassida fuscorufa species	accepted (GBIF)	No associated record in PBDB
Cassida fuscosignata species	accepted (GBIF)	No associated record in PBDB
Cassida fuscosparsa species	accepted (GBIF)	No associated record in PBDB
Cassida gansuica species	accepted (GBIF)	No associated record in PBDB
Cassida gentilis species	accepted (GBIF)	No associated record in PBDB
Cassida ghesquieri species	accepted (GBIF)	No associated record in PBDB
Cassida gilva species	accepted (GBIF)	No associated record in PBDB
Cassida ginpinica species	accepted (GBIF)	No associated record in PBDB
Cassida glabella species	accepted (GBIF)	No associated record in PBDB
Cassida glebicolor species	accepted (GBIF)	No associated record in PBDB
Cassida goudoti species	accepted (GBIF)	No associated record in PBDB
Cassida granula species	accepted (GBIF)	No associated record in PBDB
Cassida granulicollis species	accepted (GBIF)	No associated record in PBDB
Cassida guttipennis species	accepted (GBIF)	No associated record in PBDB
Cassida hablitziae species	accepted (GBIF)	No associated record in PBDB
Cassida hainanensis species	accepted (GBIF)	No associated record in PBDB
Cassida hemisphaerica species	accepted (GBIF)	No associated record in PBDB
Cassida hermione ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Cassida hexastigma species	accepted (GBIF)	No associated record in PBDB
Cassida hova species	accepted (GBIF)	No associated record in PBDB
Cassida hovacassiformis species	accepted (GBIF)	No associated record in PBDB
Cassida humeralis species	accepted (GBIF)	No associated record in PBDB
Cassida humerosa species	accepted (GBIF)	No associated record in PBDB
Cassida hyalina species	accepted (GBIF)	No associated record in PBDB
Cassida icterica species	accepted (GBIF)	No associated record in PBDB
Cassida ictericiformis species	accepted (GBIF)	No associated record in PBDB
Cassida imbecilla species	accepted (GBIF)	No associated record in PBDB
Cassida imitatrix species	accepted (GBIF)	No associated record in PBDB
Cassida immaculicollis species	accepted (GBIF)	No associated record in PBDB
Cassida imparata species	accepted (GBIF)	No associated record in PBDB
Cassida impompalis species	accepted (GBIF)	No associated record in PBDB
Cassida inaequalis species	accepted (GBIF)	No associated record in PBDB
Cassida incallida species	accepted (GBIF)	No associated record in PBDB
Cassida inciens species	accepted (GBIF)	No associated record in PBDB
Cassida inconstans species	accepted (GBIF)	No associated record in PBDB
Cassida indochinensis species	accepted (GBIF)	No associated record in PBDB
Cassida inflaccens species	accepted (GBIF)	No associated record in PBDB
Cassida inflata species	accepted (GBIF)	No associated record in PBDB
Cassida informis species	accepted (GBIF)	No associated record in PBDB
Cassida innotata species	accepted (GBIF)	No associated record in PBDB
Cassida inopinata species	accepted (GBIF)	No associated record in PBDB
Cassida inquinata species	accepted (GBIF)	No associated record in PBDB
Cassida insulana species	accepted (GBIF)	No associated record in PBDB
Cassida interemta ✝ species	accepted (GBIF) listed (PBDB)		1	28.4 Ma Oligocene	28.4 Ma Oligocene
Cassida irregularis species	accepted (GBIF)	No associated record in PBDB
Cassida irrorata species	accepted (GBIF)	No associated record in PBDB
Cassida isarogensis species	accepted (GBIF)	No associated record in PBDB
Cassida jacobsoni species	accepted (GBIF)	No associated record in PBDB
Cassida japana species	accepted (GBIF)	No associated record in PBDB
Cassida javanica species	accepted (GBIF)	No associated record in PBDB
Cassida johnsoni species	accepted (GBIF)	No associated record in PBDB
Cassida juglans species	accepted (GBIF)	No associated record in PBDB
Cassida justa species	accepted (GBIF)	No associated record in PBDB
Cassida kinabaluensis species	accepted (GBIF)	No associated record in PBDB
Cassida koreana species	accepted (GBIF)	No associated record in PBDB
Cassida kramstae ✝ species	accepted (GBIF) listed (PBDB)		1	33.9 Ma Oligocene	33.9 Ma Oligocene
Cassida kunminica species	accepted (GBIF)	No associated record in PBDB
Cassida labiatophaga species	accepted (GBIF)	No associated record in PBDB
Cassida lacrymosa species	accepted (GBIF)	No associated record in PBDB
Cassida laetabilis species	accepted (GBIF)	No associated record in PBDB
Cassida laetifica species	accepted (GBIF)	No associated record in PBDB
Cassida langeri species	accepted (GBIF)	No associated record in PBDB
Cassida laotica species	accepted (GBIF)	No associated record in PBDB
Cassida latecincta species	accepted (GBIF)	No associated record in PBDB
Cassida lateritia species	accepted (GBIF)	No associated record in PBDB
Cassida laticollis species	accepted (GBIF)	No associated record in PBDB
Cassida leucanthemi species	accepted (GBIF)	No associated record in PBDB
Cassida limpopoana species	accepted (GBIF)	No associated record in PBDB
Cassida lineola species	accepted (GBIF)	No associated record in PBDB
Cassida linnavuorii species	accepted (GBIF)	No associated record in PBDB
Cassida liquefacta species	accepted (GBIF)	No associated record in PBDB
Cassida litigiosa species	accepted (GBIF)	No associated record in PBDB
Cassida lombocensis species	accepted (GBIF)	No associated record in PBDB
Cassida lueboensis species	accepted (GBIF)	No associated record in PBDB
Cassida lunata species	accepted (GBIF)	No associated record in PBDB
Cassida luxuriosa species	accepted (GBIF)	No associated record in PBDB
Cassida luzonica species	accepted (GBIF)	No associated record in PBDB
Cassida lycii species	accepted (GBIF)	No associated record in PBDB
Cassida lyrica species	accepted (GBIF)	No associated record in PBDB
Cassida madagascarica species	accepted (GBIF)	No associated record in PBDB
Cassida magilensis species	accepted (GBIF)	No associated record in PBDB
Cassida major species	accepted (GBIF)	No associated record in PBDB
Cassida malaysiana species	accepted (GBIF)	No associated record in PBDB
Cassida mandli species	accepted (GBIF)	No associated record in PBDB
Cassida manipuria species	accepted (GBIF)	No associated record in PBDB
Cassida margaritacea species	accepted (GBIF)	No associated record in PBDB
Cassida mariaeadelheidae species	accepted (GBIF)	No associated record in PBDB
Cassida mashonensis species	accepted (GBIF)	No associated record in PBDB
Cassida medvedevi species	accepted (GBIF)	No associated record in PBDB
Cassida megapenthes ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Cassida melanophthalma species	accepted (GBIF)	No associated record in PBDB
Cassida mera species	accepted (GBIF)	No associated record in PBDB
Cassida mindanaoensis species	accepted (GBIF)	No associated record in PBDB
Cassida mishmiensis species	accepted (GBIF)	No associated record in PBDB
Cassida mongolica species	accepted (GBIF)	No associated record in PBDB
Cassida montana species	accepted (GBIF)	No associated record in PBDB
Cassida monticola species	accepted (GBIF)	No associated record in PBDB
Cassida moori species	accepted (GBIF)	No associated record in PBDB
Cassida morondaviana species	accepted (GBIF)	No associated record in PBDB
Cassida mroczkowskii species	accepted (GBIF)	No associated record in PBDB
Cassida murraea species	accepted (GBIF)	No associated record in PBDB
Cassida mysorensis species	accepted (GBIF)	No associated record in PBDB
Cassida namibiensis species	accepted (GBIF)	No associated record in PBDB
Cassida natalensis species	accepted (GBIF)	No associated record in PBDB
Cassida navicella species	accepted (GBIF)	No associated record in PBDB
Cassida nebulosa species	accepted (GBIF)	No associated record in PBDB
Cassida nepalica species	accepted (GBIF)	No associated record in PBDB
Cassida nigriventris species	accepted (GBIF)	No associated record in PBDB
Cassida nigrocastanea species	accepted (GBIF)	No associated record in PBDB
Cassida nigrodentata species	accepted (GBIF)	No associated record in PBDB
Cassida nigroflavens species	accepted (GBIF)	No associated record in PBDB
Cassida nigrogibbosa species	accepted (GBIF)	No associated record in PBDB
Cassida nigrohumeralis species	accepted (GBIF)	No associated record in PBDB
Cassida nigroramosa species	accepted (GBIF)	No associated record in PBDB
Cassida nigroscutata species	accepted (GBIF)	No associated record in PBDB
Cassida nilgirica species	accepted (GBIF)	No associated record in PBDB
Cassida nobilis species	accepted (GBIF)	No associated record in PBDB
Cassida nucula species	accepted (GBIF)	No associated record in PBDB
Cassida nuwara species	accepted (GBIF)	No associated record in PBDB
Cassida nysea species	accepted (GBIF)	No associated record in PBDB
Cassida obenbergeri species	accepted (GBIF)	No associated record in PBDB
Cassida oberlaenderi species	accepted (GBIF)	No associated record in PBDB
Cassida obtusata species	accepted (GBIF)	No associated record in PBDB
Cassida occursans species	accepted (GBIF)	No associated record in PBDB
Cassida olympica species	accepted (GBIF)	No associated record in PBDB
Cassida ovalis species	accepted (GBIF)	No associated record in PBDB
Cassida pacholatkoi species	accepted (GBIF)	No associated record in PBDB
Cassida pagana species	accepted (GBIF)	No associated record in PBDB
Cassida paiensis species	accepted (GBIF)	No associated record in PBDB
Cassida palaestina species	accepted (GBIF)	No associated record in PBDB
Cassida paliji species	accepted (GBIF)	No associated record in PBDB
Cassida pallidicollis species	accepted (GBIF)	No associated record in PBDB
Cassida pannonica species	accepted (GBIF)	No associated record in PBDB
Cassida panzeri species	accepted (GBIF)	No associated record in PBDB
Cassida parvula species	accepted (GBIF)	No associated record in PBDB
Cassida patruelis species	accepted (GBIF)	No associated record in PBDB
Cassida pauliani species	accepted (GBIF)	No associated record in PBDB
Cassida pauxilla species	accepted (GBIF)	No associated record in PBDB
Cassida pellegrini species	accepted (GBIF)	No associated record in PBDB
Cassida penelope species	accepted (GBIF)	No associated record in PBDB
Cassida pernix species	accepted (GBIF)	No associated record in PBDB
Cassida perplexa species	accepted (GBIF)	No associated record in PBDB
Cassida persica species	accepted (GBIF)	No associated record in PBDB
Cassida persicana species	accepted (GBIF)	No associated record in PBDB
Cassida pfefferi species	accepted (GBIF)	No associated record in PBDB
Cassida physodes species	accepted (GBIF)	No associated record in PBDB
Cassida piperata species	accepted (GBIF)	No associated record in PBDB
Cassida plausibilis species	accepted (GBIF)	No associated record in PBDB
Cassida plicatula species	accepted (GBIF)	No associated record in PBDB
Cassida polymeriae species	accepted (GBIF)	No associated record in PBDB
Cassida postarcuata species	accepted (GBIF)	No associated record in PBDB
Cassida praensis species	accepted (GBIF)	No associated record in PBDB
Cassida prasina species	accepted (GBIF)	No associated record in PBDB
Cassida pretiosa species	accepted (GBIF)	No associated record in PBDB
Cassida probata species	accepted (GBIF)	No associated record in PBDB
Cassida procurva species	accepted (GBIF)	No associated record in PBDB
Cassida propitia species	accepted (GBIF)	No associated record in PBDB
Cassida prospera species	accepted (GBIF)	No associated record in PBDB
Cassida pseudosyrtica species	accepted (GBIF)	No associated record in PBDB
Cassida pubescens species	accepted (GBIF)	No associated record in PBDB
Cassida pubipennis species	accepted (GBIF)	No associated record in PBDB
Cassida pudens species	accepted (GBIF)	No associated record in PBDB
Cassida pulpa species	accepted (GBIF)	No associated record in PBDB
Cassida pulvinata species	accepted (GBIF)	No associated record in PBDB
Cassida purpuricollis species	accepted (GBIF)	No associated record in PBDB
Cassida pusilla species	accepted (GBIF)	No associated record in PBDB
Cassida pusio species	accepted (GBIF)	No associated record in PBDB
Cassida pyrenaea species	accepted (GBIF)	No associated record in PBDB
Cassida quadricolorata species	accepted (GBIF)	No associated record in PBDB
Cassida quadriramosa species	accepted (GBIF)	No associated record in PBDB
Cassida quatuordecimsignata species	accepted (GBIF)	No associated record in PBDB
Cassida queenslandica species	accepted (GBIF)	No associated record in PBDB
Cassida quinaria species	accepted (GBIF)	No associated record in PBDB
Cassida quinquemaculata species	accepted (GBIF)	No associated record in PBDB
Cassida rati species	accepted (GBIF)	No associated record in PBDB
Cassida ratina species	accepted (GBIF)	No associated record in PBDB
Cassida recondita species	accepted (GBIF)	No associated record in PBDB
Cassida reitteri species	accepted (GBIF)	No associated record in PBDB
Cassida relicta species	accepted (GBIF)	No associated record in PBDB
Cassida reticulicosta species	accepted (GBIF)	No associated record in PBDB
Cassida reticulipennis species	accepted (GBIF)	No associated record in PBDB
Cassida rhodesiaca species	accepted (GBIF)	No associated record in PBDB
Cassida ribbei species	accepted (GBIF)	No associated record in PBDB
Cassida rimosa species	accepted (GBIF)	No associated record in PBDB
Cassida rothschildi species	accepted (GBIF)	No associated record in PBDB
Cassida rubiginosa species	accepted (GBIF)	No associated record in PBDB
Cassida rubripennis species	accepted (GBIF)	No associated record in PBDB
Cassida rubromaculata species	accepted (GBIF)	No associated record in PBDB
Cassida rubroornata species	accepted (GBIF)	No associated record in PBDB
Cassida rudicollis species	accepted (GBIF)	No associated record in PBDB
Cassida rufomicans species	accepted (GBIF)	No associated record in PBDB
Cassida rufovirens species	accepted (GBIF)	No associated record in PBDB
Cassida rugipennis species	accepted (GBIF)	No associated record in PBDB
Cassida ruralis species	accepted (GBIF)	No associated record in PBDB
Cassida sabahensis species	accepted (GBIF)	No associated record in PBDB
Cassida saginata species	accepted (GBIF)	No associated record in PBDB
Cassida samangana species	accepted (GBIF)	No associated record in PBDB
Cassida sanguineoguttata species	accepted (GBIF)	No associated record in PBDB
Cassida sanguinicollis species	accepted (GBIF)	No associated record in PBDB
Cassida sanguinolenta species	accepted (GBIF)	No associated record in PBDB
Cassida sanguinosa species	accepted (GBIF)	No associated record in PBDB
Cassida sappho species	accepted (GBIF)	No associated record in PBDB
Cassida sareptana species	accepted (GBIF)	No associated record in PBDB
Cassida saucia species	accepted (GBIF)	No associated record in PBDB
Cassida sauteri species	accepted (GBIF)	No associated record in PBDB
Cassida schawalleri species	accepted (GBIF)	No associated record in PBDB
Cassida schenklingi species	accepted (GBIF)	No associated record in PBDB
Cassida schoutedeni species	accepted (GBIF)	No associated record in PBDB
Cassida scymnoides species	accepted (GBIF)	No associated record in PBDB
Cassida sekerkai species	accepted (GBIF)	No associated record in PBDB
Cassida seladonia species	accepted (GBIF)	No associated record in PBDB
Cassida semipunctata species	accepted (GBIF)	No associated record in PBDB
Cassida senicula species	accepted (GBIF)	No associated record in PBDB
Cassida seniculoides species	accepted (GBIF)	No associated record in PBDB
Cassida septemdecimpunctata species	accepted (GBIF)	No associated record in PBDB
Cassida seraphina species	accepted (GBIF)	No associated record in PBDB
Cassida sexguttata species	accepted (GBIF)	No associated record in PBDB
Cassida sigillata species	accepted (GBIF)	No associated record in PBDB
Cassida signifera species	accepted (GBIF)	No associated record in PBDB
Cassida silvicola species	accepted (GBIF)	No associated record in PBDB
Cassida simanica species	accepted (GBIF)	No associated record in PBDB
Cassida smaragdocruciata species	accepted (GBIF)	No associated record in PBDB
Cassida socialis species	accepted (GBIF)	No associated record in PBDB
Cassida sodalis species	accepted (GBIF)	No associated record in PBDB
Cassida solida species	accepted (GBIF)	No associated record in PBDB
Cassida somalica species	accepted (GBIF)	No associated record in PBDB
Cassida spaethi species	accepted (GBIF)	No associated record in PBDB
Cassida spaethiana species	accepted (GBIF)	No associated record in PBDB
Cassida spartea species	accepted (GBIF)	No associated record in PBDB
Cassida spatiosa species	accepted (GBIF)	No associated record in PBDB
Cassida sphaerula species	accepted (GBIF)	No associated record in PBDB
Cassida spissa species	accepted (GBIF)	No associated record in PBDB
Cassida stevensi species	accepted (GBIF)	No associated record in PBDB
Cassida stigmatica species	accepted (GBIF)	No associated record in PBDB
Cassida strejceki species	accepted (GBIF)	No associated record in PBDB
Cassida strigaticollis species	accepted (GBIF)	No associated record in PBDB
Cassida strumosa species	accepted (GBIF)	No associated record in PBDB
Cassida stupa species	accepted (GBIF)	No associated record in PBDB
Cassida suaveola species	accepted (GBIF)	No associated record in PBDB
Cassida subacuticollis species	accepted (GBIF)	No associated record in PBDB
Cassida sublesta species	accepted (GBIF)	No associated record in PBDB
Cassida subprobata species	accepted (GBIF)	No associated record in PBDB
Cassida subreticulata species	accepted (GBIF)	No associated record in PBDB
Cassida subtilis species	accepted (GBIF)	No associated record in PBDB
Cassida successiva species	accepted (GBIF)	No associated record in PBDB
Cassida sulfurea species	accepted (GBIF)	No associated record in PBDB
Cassida sulphurago species	accepted (GBIF)	No associated record in PBDB
Cassida sulphurea species	accepted (GBIF)	No associated record in PBDB
Cassida sussamyrica species	accepted (GBIF)	No associated record in PBDB
Cassida syrtica species	accepted (GBIF)	No associated record in PBDB
Cassida taediosa species	accepted (GBIF)	No associated record in PBDB
Cassida tarda species	accepted (GBIF)	No associated record in PBDB
Cassida tenasserimensis species	accepted (GBIF)	No associated record in PBDB
Cassida tenax species	accepted (GBIF)	No associated record in PBDB
Cassida tenuicula species	accepted (GBIF)	No associated record in PBDB
Cassida thailandica species	accepted (GBIF)	No associated record in PBDB
Cassida thomsoni species	accepted (GBIF)	No associated record in PBDB
Cassida tianshanica species	accepted (GBIF)	No associated record in PBDB
Cassida timefacta species	accepted (GBIF)	No associated record in PBDB
Cassida timorensis species	accepted (GBIF)	No associated record in PBDB
Cassida tosta species	accepted (GBIF)	No associated record in PBDB
Cassida transcaucasica species	accepted (GBIF)	No associated record in PBDB
Cassida trepidula species	accepted (GBIF)	No associated record in PBDB
Cassida triangulum species	accepted (GBIF)	No associated record in PBDB
Cassida troglodytes species	accepted (GBIF)	No associated record in PBDB
Cassida trossula species	accepted (GBIF)	No associated record in PBDB
Cassida truncatipennis species	accepted (GBIF)	No associated record in PBDB
Cassida tsaratanana species	accepted (GBIF)	No associated record in PBDB
Cassida tsinlinica species	accepted (GBIF)	No associated record in PBDB
Cassida tuberculata species	accepted (GBIF)	No associated record in PBDB
Cassida tumidicollis species	accepted (GBIF)	No associated record in PBDB
Cassida turcmenica species	accepted (GBIF)	No associated record in PBDB
Cassida umbonata species	accepted (GBIF)	No associated record in PBDB
Cassida undecimnotata species	accepted (GBIF)	No associated record in PBDB
Cassida unica species	accepted (GBIF)	No associated record in PBDB
Cassida unicatenata species	accepted (GBIF)	No associated record in PBDB
Cassida unimaculata species	accepted (GBIF)	No associated record in PBDB
Cassida uniorbis species	accepted (GBIF)	No associated record in PBDB
Cassida vafra species	accepted (GBIF)	No associated record in PBDB
Cassida variabilis species	accepted (GBIF)	No associated record in PBDB
Cassida varians species	accepted (GBIF)	No associated record in PBDB
Cassida varicornis species	accepted (GBIF)	No associated record in PBDB
Cassida velaris species	accepted (GBIF)	No associated record in PBDB
Cassida verrucata species	accepted (GBIF)	No associated record in PBDB
Cassida versicolor species	accepted (GBIF)	No associated record in PBDB
Cassida vesicularis species	accepted (GBIF)	No associated record in PBDB
Cassida vespertilio species	accepted (GBIF)	No associated record in PBDB
Cassida vespertina species	accepted (GBIF)	No associated record in PBDB
Cassida vibex species	accepted (GBIF)	No associated record in PBDB
Cassida vicinalis species	accepted (GBIF)	No associated record in PBDB
Cassida vietnamica species	accepted (GBIF)	No associated record in PBDB
Cassida vigintimaculata species	accepted (GBIF)	No associated record in PBDB
Cassida virguncula species	accepted (GBIF)	No associated record in PBDB
Cassida viridiguttata species	accepted (GBIF)	No associated record in PBDB
Cassida viridinotata species	accepted (GBIF)	No associated record in PBDB
Cassida viridipennis species	accepted (GBIF)	No associated record in PBDB
Cassida viridis species	accepted (GBIF)	No associated record in PBDB
Cassida vitalisi species	accepted (GBIF)	No associated record in PBDB
Cassida vittata species	accepted (GBIF)	No associated record in PBDB
Cassida weinmanni species	accepted (GBIF)	No associated record in PBDB
Cassida weisei species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Dolichocassida Günther, 1958	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Dolichocassida*
Mionycha Weise, 1891	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Mionycha*
Cassidula Weise, 1891	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Cassidula*
Odontionycha Weise, 1891	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Odontionycha*
Cyrtonocassis Chen & Zia, 1961	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Cyrtonocassis*
Eremocassis Spaeth, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Eremocassis*
Tylocentra Spaeth & Reitter, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Tylocentra*
Crepidaspis Spaeth, 1912	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Crepidaspis*
Mionychella Spaeth, 1952	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Mionychella*
Taiwania Spaeth, 1913	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Taiwania*
Onychocassis Spaeth & Reitter, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Onychocassis*
Lordiconia Spaeth & Reitter, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Lordiconia*
Deloyala Redtenbacher, 1858	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Deloyala*
Betacassida Steinhausen, 2002	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Betacassida*
Pseudocassida Desbrochers, 1891	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Pseudocassida*
Cyclocassida Chen & Zia, 1961	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Cyclocassida*
Alledoya Hincks, 1950	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Alledoya*
Lordicassis Spaeth & Reitter, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Lordicassis*
Lasiocassis Gressitt, 1952	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Lasiocassis*
Cassidulella Strand, 1928	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Cassidulella*
Pseudocassis Steinhausen, 2002	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Pseudocassis*
Yunocassis Chen & Zia, 1961	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Chrysomelidae : *Yunocassis*

Synonymy List

Year	Name and Author
1758	Cassida Linnaeus p. 362
1761	Cassida Linnaeus p. 152
1762	Cassida Geoffroy p. 310
1763	Cassida Scopoli p. 37
1764	Cassida Müller p. 5
1766	Cassida Schaeffer
1767	Cassida Linnaeus p. 574

Year	Name and Author
1775	Cassida Fabricius p. 88
1776	Cassida Müller p. 65
1796	Cassida Latreille p. 62
1798	Cassida Fabricius p. 81
1802	Cassida Latreille p. 233
2011	Cassida Bouchard et al. p. 512

References

Linnaeus C. , Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Decima 1, 1-824 doi:10.5962/bhl.title.542

Linnaeus C. , Fauna Svecica: Sistens Animalia Sveciae Regni: Mammalia, Aves, Amphibia, Pisces, Insecta, Vermes, Distributa per Classes & Ordines, Genera & Species, cum Differentiis Specierum, Synonymis Auctorum, Nominibus Incolarum, Locis Natalium, Descriptionibus Insectorum, 1-578

Geoffroy E. L. , Histoire Abregée des Insectes qui se Trouvent aux Environs de Paris, Tome Premier, 1-523

Scopoli G. A. , Entomologica Carniolica Exhibens Insecta Carnioliae Indigena et Distributa in Ordines, Genera, Species, Varietates Methodo Linnaeana, 1-421

Müller O. F. , Fauna Insectorum Fridrichsdalina, Sive Methodica Descriptio Insectorum Agri Fridrichsdalensis, cum Characteribus Genericis et Specificis, Nominibus Trivialibus, Locis Natalibus, Iconibus Allegatis, Novisque Pluribus Speciebus Additis, 1-96

Linnaeus C. , Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. Editio Duodecima 1 2, 533-1327

Fabricius J. C. , Systema Entomologiae, Sistens Insectorum Classes, Ordines, Genera, Species Adiectis Synonymis, Locis, Descriptionibus, Observationibus, 1-832

Müller O. F. , Zoologicae Danicae Prodromus, seu Animalium Daniae et Norvegiae Indigenarum. Characteres, Nomina et Synonyma Imprimis Popularium, 1-281

Latreille P. A. , Précis des Caractères Génériques des Insectes Disposés dans un Ordre Naturel, 1-201

Fabricius J. C. , Supplementum Entomologiae Systematicae, 1-572

Latreille P. A. , Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième, 1-467

Bouchard P., Bousquet Y., et al (2011) Family-group names in Coleoptera (Insecta), ZooKeys 88, 1-972

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

SN2000: Brands, S. J. (compiler) 1989-2005. Systema Naturae 2000. Amsterdam, The Netherlands (2006 version). Available online at http://sn2000.taxonomy.nl/. - via The Interim Register of Marine and Nonmarine Genera

WoRMS (Mar 2013) - via The Interim Register of Marine and Nonmarine Genera

as per family - via The Interim Register of Marine and Nonmarine Genera

International Commission on Zoological Nomenclature. Opinion 1023 Cassidae (Mollusca) and Cassidinae (Insecta): placed on the Official List of Family-group Names in Zoology.Bulletin of Zoological Nomenclature 31, 127-129 (1974) - via Official Lists and Indexes of Names in Zoology

Linnaeus, C. (1758). Systema Naturae per Regna Tria Naturae, Secundum Classes, Ordines, Genera, Species, cum Characteribus, Differentiis, Synonymis, Locis. 10th edition. - via The Interim Register of Marine and Nonmarine Genera

Riley EG (1986b) Notes on Cassida relicta, a tortoise beetle endemic to North America, with a key to Nearctic species of Cassida (Coleoptera: Chrysomelidae). Entomological News 97: 141-146. - via Checklist of Beetles (Coleoptera) of Canada and Alaska. Second Edition.

Hayward, P.J.; Ryland, J.S. (Ed.). (1990). The marine fauna of the British Isles and North-West Europe: 1. Introduction and protozoans to arthropods. Clarendon Press: Oxford, UK. ISBN 0-19-857356-1. 627 pp. - via World Register of Marine Species

Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

Borowiec, Lech, 1999: null. A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). 476. - via Integrated Taxonomic Information System (ITIS)

Riley, Edward G., Shawn M. Clark, and Terry N. Seeno, 2003: Catalog of the leaf beetles of America north of Mexico (Coleoptera: Megalopodidae, Orsodacnidae and Chrysomelidae, excluding Bruchinae). Coleopterists Society Special Publication no. 1. 290. - via Integrated Taxonomic Information System (ITIS)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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