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Harpalus

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Description

Harpalus (Greek: Ἅρπαλος) son of Machatas was an aristocrat of Macedon and boyhood friend of Alexander the Great in the 4th century BC. Being lame in a leg, and therefore exempt from military service, Harpalus did not follow Alexander in his advance within the Persian Empire but received nonetheless a post in Asia Minor. Alexander reportedly contacted him with a demand of reading material for his spare time. Harpalus sent his King theatrical plays byAeschylus, Sophocles and Euripides, the history of Philistus and odes by Philoxenus and Telestes.

From Wikipedia article at https://en.wikipedia.org/wiki/Harpalus, which is released under the Creative Commons Attribution-Share-Alike License 3.0.

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	1035821	https://www.gbif.org/species/1035821
PaleoBioDB ID (PBDB)	196066	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=196066

Rank

genus

Taxonomy (GBIF,PBDB)

Life : Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : Harpalus

Taxonomic Status (GBIF)

accepted

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Bilateria
-	Eubilateria	Ax 1987
-	Protostomia	Grobben 1908
-	Ecdysozoa
-	Panarthropoda
phylum	Arthropoda	Latreille 1829
subphylum	Mandibulata
-	Pancrustacea
superclass	Hexapoda	Latreille 1825
-	Cercofilata	Kukalová-Peck 1987
class	Insecta	Linnaeus 1758
-	Dicondylia
-	Paranotalia	Sroka et al. 2014
subclass	Pterygota	Brauer 1885
-	Neopterygota	Crampton 1924
infraclass	Neoptera	Martynov 1923
-	Holometabola
-	Aparaglossata
-	Coleopterida	Kukalová-Peck and Lawrence 2004
order	Coleoptera	Linnaeus 1758
suborder	Adephaga	Clairville 1806
family	Carabidae	Latreille 1802
subfamily	Harpalinae
tribe	Harpalini	Bonelli 1810
subtribe	Harpalina	Bonelli 1810
genus	Harpalus	Latreille 1802

Scientific Name

Harpalus Latreille, 1802

Name Published In

Latreille, Pierre A. 1802. Histoire naturelle, générale et particulière des crustacés et des insectes. rédigé par C. S. Sonnini. F. Dufart, Paris. Vol. 3: i-xii; 13-467.

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Harpalus	genus	belongs to Melanchlaeni	stated with evidence	Latreille, 1802
Harpalus	genus	belongs to Carabiques	stated with evidence	Dejean, 1821
Harpalus	genus	belongs to Harpali	stated with evidence	LeConte, 1861
Harpalus	genus	belongs to Carabidae	stated with evidence	Scudder, 1900
Harpalus	genus	belongs to Carabidae	stated without evidence	Cockerell, 1921
Harpalus	genus	belongs to Harpalina	stated with evidence	Bouchard et al., 2011
Harpalus	genus	belongs to Harpalini	stated without evidence	Tamutis et al., 2011

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

4 (14%)

First Recorded Appearance

66.0 - 59.2 Ma
Paleocene

Environment

terrestrial (based on Insecta)

Motility

actively mobile (based on Arthropoda)

Taphonomy

chitin (based on Insecta)

Primary Reference (PBDB)

P. A. Latreille. 1802. Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième

Wikipedia

https://en.wikipedia.org/wiki/Harpalus

External Images

Fossil Distribution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Harpalus abolitus ✝ species	accepted (GBIF) listed (PBDB)		1	28.4 Ma Oligocene	28.4 Ma Oligocene
Harpalus actiosus species	accepted (GBIF)	No associated record in PBDB
Harpalus acupalpoides species	accepted (GBIF)	No associated record in PBDB
Harpalus adenticulatus species	accepted (GBIF)	No associated record in PBDB
Harpalus aeneipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus aenigma species	accepted (GBIF)	No associated record in PBDB
Harpalus aequicollis species	accepted (GBIF)	No associated record in PBDB
Harpalus affinis species	accepted (GBIF)	No associated record in PBDB
Harpalus agakhaniantzi species	accepted (GBIF)	No associated record in PBDB
Harpalus agnatus species	accepted (GBIF)	No associated record in PBDB
Harpalus akinini species	accepted (GBIF)	No associated record in PBDB
Harpalus alajensis species	accepted (GBIF)	No associated record in PBDB
Harpalus albanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus alexandrae species	accepted (GBIF)	No associated record in PBDB
Harpalus alexeevi species	accepted (GBIF)	No associated record in PBDB
Harpalus alienus species	accepted (GBIF)	No associated record in PBDB
Harpalus alpivagus species	accepted (GBIF)	No associated record in PBDB
Harpalus amarellus species	accepted (GBIF)	No associated record in PBDB
Harpalus amariformis species	accepted (GBIF)	No associated record in PBDB
Harpalus amplicollis species	accepted (GBIF)	No associated record in PBDB
Harpalus amputatus species	accepted (GBIF) listed (PBDB)		16	2.59 Ma Pleistocene	0 Ma Extant
Harpalus anactus species	accepted (GBIF)	No associated record in PBDB
Harpalus anatolicus species	accepted (GBIF)	No associated record in PBDB
Harpalus angulatus species	accepted (GBIF)	No associated record in PBDB
Harpalus angustipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus angustitarsis species	accepted (GBIF)	No associated record in PBDB
Harpalus animosus species	accepted (GBIF)	No associated record in PBDB
Harpalus anisodactyliformis species	accepted (GBIF)	No associated record in PBDB
Harpalus antonowi species	accepted (GBIF)	No associated record in PBDB
Harpalus anxioides species	accepted (GBIF)	No associated record in PBDB
Harpalus anxius species	accepted (GBIF)	No associated record in PBDB
Harpalus aogashimensis species	accepted (GBIF)	No associated record in PBDB
Harpalus apache species	accepted (GBIF)	No associated record in PBDB
Harpalus araraticus species	accepted (GBIF)	No associated record in PBDB
Harpalus arcuatus species	accepted (GBIF)	No associated record in PBDB
Harpalus arnoldii species	accepted (GBIF)	No associated record in PBDB
Harpalus ascetes species	accepted (GBIF)	No associated record in PBDB
Harpalus asemus species	accepted (GBIF)	No associated record in PBDB
Harpalus aterrimus species	accepted (GBIF)	No associated record in PBDB
Harpalus atratus species	accepted (GBIF)	No associated record in PBDB
Harpalus atrichatus species	accepted (GBIF)	No associated record in PBDB
Harpalus atripes species	accepted (GBIF)	No associated record in PBDB
Harpalus attenuatus species	accepted (GBIF)	No associated record in PBDB
Harpalus australasiae species	accepted (GBIF)	No associated record in PBDB
Harpalus autumnalis species	accepted (GBIF)	No associated record in PBDB
Harpalus azumai species	accepted (GBIF)	No associated record in PBDB
Harpalus babai species	accepted (GBIF)	No associated record in PBDB
Harpalus badakschanus species	accepted (GBIF)	No associated record in PBDB
Harpalus baleensis species	accepted (GBIF)	No associated record in PBDB
Harpalus balli species	accepted (GBIF)	No associated record in PBDB
Harpalus basanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus basilewskyi species	accepted (GBIF)	No associated record in PBDB
Harpalus basuto species	accepted (GBIF)	No associated record in PBDB
Harpalus bellieri species	accepted (GBIF)	No associated record in PBDB
Harpalus beneshi species	accepted (GBIF)	No associated record in PBDB
Harpalus bitinctus species	accepted (GBIF)	No associated record in PBDB
Harpalus brachypterus species	accepted (GBIF)	No associated record in PBDB
Harpalus brevicornis species	accepted (GBIF)	No associated record in PBDB
Harpalus brevis species	accepted (GBIF)	No associated record in PBDB
Harpalus breviusculus species	accepted (GBIF)	No associated record in PBDB
Harpalus bruckmanni ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus brunnipes species	accepted (GBIF)	No associated record in PBDB
Harpalus bucharicus species	accepted (GBIF)	No associated record in PBDB
Harpalus bungii species	accepted (GBIF)	No associated record in PBDB
Harpalus burmeisteri species	accepted (GBIF)	No associated record in PBDB
Harpalus caeruleatus species	accepted (GBIF)	No associated record in PBDB
Harpalus caiphus species	accepted (GBIF)	No associated record in PBDB
Harpalus calathoides species	accepted (GBIF)	No associated record in PBDB
Harpalus calceatus species	accepted (GBIF)	No associated record in PBDB
Harpalus caliginosus species	accepted (GBIF)	No associated record in PBDB
Harpalus capicola species	accepted (GBIF)	No associated record in PBDB
Harpalus capito species	accepted (GBIF)	No associated record in PBDB
Harpalus cardoni species	accepted (GBIF)	No associated record in PBDB
Harpalus caspius species	accepted (GBIF)	No associated record in PBDB
Harpalus cautus species	accepted (GBIF)	No associated record in PBDB
Harpalus celax species	accepted (GBIF)	No associated record in PBDB
Harpalus cephalotes species	accepted (GBIF)	No associated record in PBDB
Harpalus chalcentus species	accepted (GBIF)	No associated record in PBDB
Harpalus chalcopterus species	accepted (GBIF)	No associated record in PBDB
Harpalus chasanensis species	accepted (GBIF)	No associated record in PBDB
Harpalus chengjiangensis species	accepted (GBIF)	No associated record in PBDB
Harpalus chiloschizontus species	accepted (GBIF)	No associated record in PBDB
Harpalus chobautianus species	accepted (GBIF)	No associated record in PBDB
Harpalus chrysopus species	accepted (GBIF)	No associated record in PBDB
Harpalus cilihumerus species	accepted (GBIF)	No associated record in PBDB
Harpalus circumpunctatus species	accepted (GBIF)	No associated record in PBDB
Harpalus cisteloides species	accepted (GBIF)	No associated record in PBDB
Harpalus cohni species	accepted (GBIF)	No associated record in PBDB
Harpalus colchicus species	accepted (GBIF)	No associated record in PBDB
Harpalus compar species	accepted (GBIF)	No associated record in PBDB
Harpalus compressus species	accepted (GBIF)	No associated record in PBDB
Harpalus conditus ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Harpalus constrictus ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus contemptus species	accepted (GBIF)	No associated record in PBDB
Harpalus convexus species	accepted (GBIF)	No associated record in PBDB
Harpalus cordifer species	accepted (GBIF)	No associated record in PBDB
Harpalus coreanus species	accepted (GBIF)	No associated record in PBDB
Harpalus corporosus species	accepted (GBIF)	No associated record in PBDB
Harpalus corpulentus species	accepted (GBIF)	No associated record in PBDB
Harpalus corrugatus species	accepted (GBIF)	No associated record in PBDB
Harpalus crates species	accepted (GBIF)	No associated record in PBDB
Harpalus cupreus species	accepted (GBIF)	No associated record in PBDB
Harpalus cursorius species	accepted (GBIF)	No associated record in PBDB
Harpalus cyanopterus species	accepted (GBIF)	No associated record in PBDB
Harpalus cyclogonus species	accepted (GBIF)	No associated record in PBDB
Harpalus cyrenaicus species	accepted (GBIF)	No associated record in PBDB
Harpalus danieli species	accepted (GBIF)	No associated record in PBDB
Harpalus davidi species	accepted (GBIF)	No associated record in PBDB
Harpalus davidianus species	accepted (GBIF)	No associated record in PBDB
Harpalus decipiens species	accepted (GBIF)	No associated record in PBDB
Harpalus defector species	accepted (GBIF)	No associated record in PBDB
Harpalus deletus ✝ species	accepted (GBIF) listed (PBDB)		1	28.1 Ma Oligocene	28.1 Ma Oligocene
Harpalus demelti species	accepted (GBIF)	No associated record in PBDB
Harpalus desertus species	accepted (GBIF)	No associated record in PBDB
Harpalus dilatatus ✝ species	accepted (GBIF) listed (PBDB)		1	33.9 Ma Oligocene	33.9 Ma Oligocene
Harpalus diligens species	accepted (GBIF)	No associated record in PBDB
Harpalus dimidiatus species	accepted (GBIF)	No associated record in PBDB
Harpalus disaogashimensis species	accepted (GBIF)	No associated record in PBDB
Harpalus disimuciulus species	accepted (GBIF)	No associated record in PBDB
Harpalus dispar species	accepted (GBIF)	No associated record in PBDB
Harpalus dissitus species	accepted (GBIF)	No associated record in PBDB
Harpalus distinguendus species	accepted (GBIF)	No associated record in PBDB
Harpalus diversicollis species	accepted (GBIF)	No associated record in PBDB
Harpalus dubius species	accepted (GBIF)	No associated record in PBDB
Harpalus durangoensis species	accepted (GBIF)	No associated record in PBDB
Harpalus ebeninus species	accepted (GBIF)	No associated record in PBDB
Harpalus egenus species	accepted (GBIF)	No associated record in PBDB
Harpalus egorovi species	accepted (GBIF)	No associated record in PBDB
Harpalus ellipsis species	accepted (GBIF)	No associated record in PBDB
Harpalus eous species	accepted (GBIF)	No associated record in PBDB
Harpalus erosus species	accepted (GBIF)	No associated record in PBDB
Harpalus erraticus species	accepted (GBIF)	No associated record in PBDB
Harpalus erythropus species	accepted (GBIF)	No associated record in PBDB
Harpalus euchlorus species	accepted (GBIF)	No associated record in PBDB
Harpalus ewaldi species	accepted (GBIF)	No associated record in PBDB
Harpalus excavatus ✝ species	accepted (GBIF) listed (PBDB)		1	33.9 Ma Oligocene	33.9 Ma Oligocene
Harpalus exiguus species	accepted (GBIF)	No associated record in PBDB
Harpalus famelicus species	accepted (GBIF)	No associated record in PBDB
Harpalus farkaci species	accepted (GBIF)	No associated record in PBDB
Harpalus faunus species	accepted (GBIF)	No associated record in PBDB
Harpalus ferghanensis species	accepted (GBIF)	No associated record in PBDB
Harpalus fimetarius species	accepted (GBIF)	No associated record in PBDB
Harpalus flavescens species	accepted (GBIF)	No associated record in PBDB
Harpalus flavicornis species	accepted (GBIF)	No associated record in PBDB
Harpalus fokienensis species	accepted (GBIF)	No associated record in PBDB
Harpalus forsstroemi species	accepted (GBIF)	No associated record in PBDB
Harpalus foveiger species	accepted (GBIF)	No associated record in PBDB
Harpalus franzi species	accepted (GBIF)	No associated record in PBDB
Harpalus frater species	accepted (GBIF)	No associated record in PBDB
Harpalus fraternus species	accepted (GBIF)	No associated record in PBDB
Harpalus froelichii species	accepted (GBIF)	No associated record in PBDB
Harpalus fukiensis species	accepted (GBIF)	No associated record in PBDB
Harpalus fulgens species	accepted (GBIF)	No associated record in PBDB
Harpalus fulvicornis species	accepted (GBIF)	No associated record in PBDB
Harpalus fulvilabris species	accepted (GBIF)	No associated record in PBDB
Harpalus fulvipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus fulvus species	accepted (GBIF)	No associated record in PBDB
Harpalus fuscicornis species	accepted (GBIF)	No associated record in PBDB
Harpalus fuscipalpis species	accepted (GBIF)	No associated record in PBDB
Harpalus fuscipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus fuscoaeneus species	accepted (GBIF)	No associated record in PBDB
Harpalus ganssuensis species	accepted (GBIF)	No associated record in PBDB
Harpalus geniculatus species	accepted (GBIF)	No associated record in PBDB
Harpalus giacomazzoi species	accepted (GBIF)	No associated record in PBDB
Harpalus gilgil species	accepted (GBIF)	No associated record in PBDB
Harpalus gisellae species	accepted (GBIF)	No associated record in PBDB
Harpalus glasunovi species	accepted (GBIF)	No associated record in PBDB
Harpalus gravis species	accepted (GBIF)	No associated record in PBDB
Harpalus gregoryi species	accepted (GBIF)	No associated record in PBDB
Harpalus grilli species	accepted (GBIF)	No associated record in PBDB
Harpalus griseus species	accepted (GBIF)	No associated record in PBDB
Harpalus hartmanni species	accepted (GBIF)	No associated record in PBDB
Harpalus hatchi species	accepted (GBIF)	No associated record in PBDB
Harpalus hauserianus species	accepted (GBIF)	No associated record in PBDB
Harpalus heeri species	accepted (GBIF)	No associated record in PBDB
Harpalus herbivagus species	accepted (GBIF)	No associated record in PBDB
Harpalus heyrovskyi species	accepted (GBIF)	No associated record in PBDB
Harpalus hirtipes species	accepted (GBIF)	No associated record in PBDB
Harpalus honestus species	accepted (GBIF)	No associated record in PBDB
Harpalus hospes species	accepted (GBIF)	No associated record in PBDB
Harpalus huachuca species	accepted (GBIF)	No associated record in PBDB
Harpalus hybridus species	accepted (GBIF)	No associated record in PBDB
Harpalus hypogeomysis species	accepted (GBIF)	No associated record in PBDB
Harpalus idiotus species	accepted (GBIF)	No associated record in PBDB
Harpalus imerinae species	accepted (GBIF)	No associated record in PBDB
Harpalus immixtus species	accepted (GBIF)	No associated record in PBDB
Harpalus impressicollis species	accepted (GBIF)	No associated record in PBDB
Harpalus impressus species	accepted (GBIF)	No associated record in PBDB
Harpalus inconcinnus species	accepted (GBIF)	No associated record in PBDB
Harpalus indianus species	accepted (GBIF)	No associated record in PBDB
Harpalus indicola species	accepted (GBIF)	No associated record in PBDB
Harpalus indicus species	accepted (GBIF)	No associated record in PBDB
Harpalus indigens species	accepted (GBIF)	No associated record in PBDB
Harpalus inexpectatus species	accepted (GBIF)	No associated record in PBDB
Harpalus inexspectatus species	accepted (GBIF)	No associated record in PBDB
Harpalus ingenuus species	accepted (GBIF)	No associated record in PBDB
Harpalus innocuus species	accepted (GBIF)	No associated record in PBDB
Harpalus italus species	accepted (GBIF)	No associated record in PBDB
Harpalus janinae species	accepted (GBIF)	No associated record in PBDB
Harpalus jeanneli species	accepted (GBIF)	No associated record in PBDB
Harpalus jordanus species	accepted (GBIF)	No associated record in PBDB
Harpalus jureceki species	accepted (GBIF)	No associated record in PBDB
Harpalus kabakianus species	accepted (GBIF)	No associated record in PBDB
Harpalus kabakovi species	accepted (GBIF)	No associated record in PBDB
Harpalus kadleci species	accepted (GBIF)	No associated record in PBDB
Harpalus kadyrbekovi species	accepted (GBIF)	No associated record in PBDB
Harpalus kagyzmanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus kailiensis species	accepted (GBIF)	No associated record in PBDB
Harpalus kandaharensis species	accepted (GBIF)	No associated record in PBDB
Harpalus karakorum species	accepted (GBIF)	No associated record in PBDB
Harpalus karamani species	accepted (GBIF)	No associated record in PBDB
Harpalus katiae species	accepted (GBIF)	No associated record in PBDB
Harpalus kazanensis species	accepted (GBIF)	No associated record in PBDB
Harpalus kaznakovi species	accepted (GBIF)	No associated record in PBDB
Harpalus kibonoti species	accepted (GBIF)	No associated record in PBDB
Harpalus kirgisicus species	accepted (GBIF)	No associated record in PBDB
Harpalus kiritshenkoi species	accepted (GBIF)	No associated record in PBDB
Harpalus klapperichi species	accepted (GBIF)	No associated record in PBDB
Harpalus kmecoi species	accepted (GBIF)	No associated record in PBDB
Harpalus knorri species	accepted (GBIF)	No associated record in PBDB
Harpalus koshiensis species	accepted (GBIF)	No associated record in PBDB
Harpalus kozlovi species	accepted (GBIF)	No associated record in PBDB
Harpalus kryzhanovskii species	accepted (GBIF)	No associated record in PBDB
Harpalus kunarensis species	accepted (GBIF)	No associated record in PBDB
Harpalus laetus species	accepted (GBIF)	No associated record in PBDB
Harpalus laevipes species	accepted (GBIF)	No associated record in PBDB
Harpalus lama species	accepted (GBIF)	No associated record in PBDB
Harpalus lampronotus species	accepted (GBIF)	No associated record in PBDB
Harpalus lateralis species	accepted (GBIF)	No associated record in PBDB
Harpalus laticeps species	accepted (GBIF)	No associated record in PBDB
Harpalus latus species	accepted (GBIF)	No associated record in PBDB
Harpalus lederi species	accepted (GBIF)	No associated record in PBDB
Harpalus lethierryi species	accepted (GBIF)	No associated record in PBDB
Harpalus lewisii species	accepted (GBIF)	No associated record in PBDB
Harpalus lheritieri ✝ species	accepted (GBIF) listed (PBDB)		1	59.2 Ma Paleocene	59.2 Ma Paleocene
Harpalus liasinus species	accepted (GBIF)	No associated record in PBDB
Harpalus liobasis species	accepted (GBIF)	No associated record in PBDB
Harpalus litigiosus species	accepted (GBIF)	No associated record in PBDB
Harpalus longihornus species	accepted (GBIF)	No associated record in PBDB
Harpalus longipalmatus species	accepted (GBIF)	No associated record in PBDB
Harpalus lopezi species	accepted (GBIF)	No associated record in PBDB
Harpalus lugubris species	accepted (GBIF)	No associated record in PBDB
Harpalus lumbaris species	accepted (GBIF)	No associated record in PBDB
Harpalus luteicornis species	accepted (GBIF)	No associated record in PBDB
Harpalus lutshniki species	accepted (GBIF)	No associated record in PBDB
Harpalus maceratus ✝ species	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Harpalus macronotus species	accepted (GBIF)	No associated record in PBDB
Harpalus madagascariensis species	accepted (GBIF)	No associated record in PBDB
Harpalus mairei species	accepted (GBIF)	No associated record in PBDB
Harpalus major species	accepted (GBIF)	No associated record in PBDB
Harpalus makhekensis species	accepted (GBIF)	No associated record in PBDB
Harpalus manas species	accepted (GBIF)	No associated record in PBDB
Harpalus marginellus species	accepted (GBIF)	No associated record in PBDB
Harpalus mariae species	accepted (GBIF)	No associated record in PBDB
Harpalus martini species	accepted (GBIF)	No associated record in PBDB
Harpalus masoreoides species	accepted (GBIF)	No associated record in PBDB
Harpalus massarti species	accepted (GBIF)	No associated record in PBDB
Harpalus mauritanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus medvedevi species	accepted (GBIF)	No associated record in PBDB
Harpalus megacephalus species	accepted (GBIF)	No associated record in PBDB
Harpalus meghalayensis species	accepted (GBIF)	No associated record in PBDB
Harpalus melancholicus species	accepted (GBIF)	No associated record in PBDB
Harpalus melaneus species	accepted (GBIF)	No associated record in PBDB
Harpalus meridianus species	accepted (GBIF)	No associated record in PBDB
Harpalus metallinus species	accepted (GBIF)	No associated record in PBDB
Harpalus metarsius species	accepted (GBIF)	No associated record in PBDB
Harpalus meteorus species	accepted (GBIF)	No associated record in PBDB
Harpalus michaili species	accepted (GBIF)	No associated record in PBDB
Harpalus michailovi species	accepted (GBIF)	No associated record in PBDB
Harpalus microdemas species	accepted (GBIF)	No associated record in PBDB
Harpalus microthorax species	accepted (GBIF)	No associated record in PBDB
Harpalus miles species	accepted (GBIF)	No associated record in PBDB
Harpalus minutulus species	accepted (GBIF)	No associated record in PBDB
Harpalus mitridati species	accepted (GBIF)	No associated record in PBDB
Harpalus mlynari species	accepted (GBIF)	No associated record in PBDB
Harpalus modestus species	accepted (GBIF)	No associated record in PBDB
Harpalus morvani species	accepted (GBIF)	No associated record in PBDB
Harpalus muciulus species	accepted (GBIF)	No associated record in PBDB
Harpalus nanniscus species	accepted (GBIF)	No associated record in PBDB
Harpalus natalensis species	accepted (GBIF)	No associated record in PBDB
Harpalus natalicus species	accepted (GBIF)	No associated record in PBDB
Harpalus neglectus species	accepted (GBIF)	No associated record in PBDB
Harpalus nero ✝ species	accepted (GBIF) listed (PBDB)		4	28.4 Ma Oligocene	28.1 Ma Oligocene
Harpalus nevadensis species	accepted (GBIF)	No associated record in PBDB
Harpalus nigrans species	accepted (GBIF)	No associated record in PBDB
Harpalus nigritarsis species	accepted (GBIF)	No associated record in PBDB
Harpalus nitens ✝ species	accepted (GBIF) listed (PBDB)		1	66.0 Ma Late/Upper Cretaceous	66.0 Ma Late/Upper Cretaceous
Harpalus numidicus species	accepted (GBIF)	No associated record in PBDB
Harpalus nuperus ✝ species	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Harpalus nyassicus species	accepted (GBIF)	No associated record in PBDB
Harpalus obliquus species	accepted (GBIF)	No associated record in PBDB
Harpalus oblitus species	accepted (GBIF)	No associated record in PBDB
Harpalus obnixus species	accepted (GBIF)	No associated record in PBDB
Harpalus obtusiusculus species	accepted (GBIF)	No associated record in PBDB
Harpalus obtusus ✝ species	accepted (GBIF) listed (PBDB)		2	33.9 Ma Oligocene	33.9 Ma Oligocene
Harpalus ochropus species	accepted (GBIF)	No associated record in PBDB
Harpalus oodioides species	accepted (GBIF)	No associated record in PBDB
Harpalus opacipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus optabilis species	accepted (GBIF)	No associated record in PBDB
Harpalus oustaleti ✝ species	accepted (GBIF) listed (PBDB)		3	38.0 Ma Eocene	28.1 Ma Oligocene
Harpalus ovtshinnikovi species	accepted (GBIF)	No associated record in PBDB
Harpalus pachys species	accepted (GBIF)	No associated record in PBDB
Harpalus pallidipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus pallipes species	accepted (GBIF)	No associated record in PBDB
Harpalus parasinuatus species	accepted (GBIF)	No associated record in PBDB
Harpalus paratus species	accepted (GBIF)	No associated record in PBDB
Harpalus parvulus species	accepted (GBIF)	No associated record in PBDB
Harpalus pastor species	accepted (GBIF)	No associated record in PBDB
Harpalus pecinai species	accepted (GBIF)	No associated record in PBDB
Harpalus penglainus species	accepted (GBIF)	No associated record in PBDB
Harpalus pensylvanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus periglabellus species	accepted (GBIF)	No associated record in PBDB
Harpalus petri species	accepted (GBIF)	No associated record in PBDB
Harpalus picipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus pilosus species	accepted (GBIF)	No associated record in PBDB
Harpalus plancyi species	accepted (GBIF)	No associated record in PBDB
Harpalus platynotus species	accepted (GBIF)	No associated record in PBDB
Harpalus pleistocenicus ✝ species	accepted (GBIF) listed (PBDB)		1	0.13 Ma Pleistocene	0.13 Ma Pleistocene
Harpalus plenalis species	accepted (GBIF)	No associated record in PBDB
Harpalus politus species	accepted (GBIF)	No associated record in PBDB
Harpalus polyglyptus species	accepted (GBIF)	No associated record in PBDB
Harpalus poncei species	accepted (GBIF)	No associated record in PBDB
Harpalus potanini species	accepted (GBIF)	No associated record in PBDB
Harpalus poussereaui species	accepted (GBIF)	No associated record in PBDB
Harpalus praecurrens species	accepted (GBIF)	No associated record in PBDB
Harpalus praticola species	accepted (GBIF)	No associated record in PBDB
Harpalus procognatus species	accepted (GBIF)	No associated record in PBDB
Harpalus progrediens species	accepted (GBIF)	No associated record in PBDB
Harpalus prosperus species	accepted (GBIF)	No associated record in PBDB
Harpalus protractus species	accepted (GBIF)	No associated record in PBDB
Harpalus providens species	accepted (GBIF)	No associated record in PBDB
Harpalus pseudohauserianus species	accepted (GBIF)	No associated record in PBDB
Harpalus pseudophonoides species	accepted (GBIF)	No associated record in PBDB
Harpalus pseudoserripes species	accepted (GBIF)	No associated record in PBDB
Harpalus pseudotinctulus species	accepted (GBIF)	No associated record in PBDB
Harpalus pterostichus species	accepted (GBIF)	No associated record in PBDB
Harpalus puetzi species	accepted (GBIF)	No associated record in PBDB
Harpalus pulchrinulus species	accepted (GBIF)	No associated record in PBDB
Harpalus pulvinatus species	accepted (GBIF)	No associated record in PBDB
Harpalus pumilus species	accepted (GBIF)	No associated record in PBDB
Harpalus punctatostriatus species	accepted (GBIF)	No associated record in PBDB
Harpalus puncticeps species	accepted (GBIF)	No associated record in PBDB
Harpalus punctipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus pusillus species	accepted (GBIF) listed (PBDB)		4	0.13 Ma Pleistocene	0 Ma Extant
Harpalus pygmaeus species	accepted (GBIF)	No associated record in PBDB
Harpalus quadratus species	accepted (GBIF)	No associated record in PBDB
Harpalus quenselii species	accepted (GBIF)	No associated record in PBDB
Harpalus raphaili species	accepted (GBIF)	No associated record in PBDB
Harpalus redivivus ✝ species	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Harpalus reflexus species	accepted (GBIF)	No associated record in PBDB
Harpalus reitteri species	accepted (GBIF)	No associated record in PBDB
Harpalus remboides species	accepted (GBIF)	No associated record in PBDB
Harpalus retractus species	accepted (GBIF)	No associated record in PBDB
Harpalus reversus species	accepted (GBIF)	No associated record in PBDB
Harpalus rivalsi species	accepted (GBIF)	No associated record in PBDB
Harpalus roninus species	accepted (GBIF)	No associated record in PBDB
Harpalus rotundus species	accepted (GBIF)	No associated record in PBDB
Harpalus rougemonti species	accepted (GBIF)	No associated record in PBDB
Harpalus rubefactus species	accepted (GBIF)	No associated record in PBDB
Harpalus rubripes species	accepted (GBIF)	No associated record in PBDB
Harpalus rufipalpis species	accepted (GBIF)	No associated record in PBDB
Harpalus rufipes species	accepted (GBIF)	No associated record in PBDB
Harpalus rufiscapus species	accepted (GBIF)	No associated record in PBDB
Harpalus rufitarsoides species	accepted (GBIF)	No associated record in PBDB
Harpalus rufomarginatus species	accepted (GBIF)	No associated record in PBDB
Harpalus rumelicus species	accepted (GBIF)	No associated record in PBDB
Harpalus salinulus species	accepted (GBIF)	No associated record in PBDB
Harpalus salinus species	accepted (GBIF)	No associated record in PBDB
Harpalus sanctaehelenae species	accepted (GBIF)	No associated record in PBDB
Harpalus saxicola species	accepted (GBIF)	No associated record in PBDB
Harpalus scabripectus species	accepted (GBIF)	No associated record in PBDB
Harpalus schaumii species	accepted (GBIF)	No associated record in PBDB
Harpalus schlotheimi species	accepted (GBIF)	No associated record in PBDB
Harpalus semenowi species	accepted (GBIF)	No associated record in PBDB
Harpalus semipunctatus species	accepted (GBIF)	No associated record in PBDB
Harpalus sericatus species	accepted (GBIF)	No associated record in PBDB
Harpalus serripes species	accepted (GBIF)	No associated record in PBDB
Harpalus servus species	accepted (GBIF)	No associated record in PBDB
Harpalus setiporus species	accepted (GBIF)	No associated record in PBDB
Harpalus seyrigi species	accepted (GBIF)	No associated record in PBDB
Harpalus siculus species	accepted (GBIF)	No associated record in PBDB
Harpalus signaticornis species	accepted (GBIF)	No associated record in PBDB
Harpalus silipes species	accepted (GBIF)	No associated record in PBDB
Harpalus simplicidens species	accepted (GBIF)	No associated record in PBDB
Harpalus singularis species	accepted (GBIF)	No associated record in PBDB
Harpalus sinicus species	accepted (GBIF)	No associated record in PBDB
Harpalus sinis ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus sinuatipennis species	accepted (GBIF)	No associated record in PBDB
Harpalus sinuatus species	accepted (GBIF)	No associated record in PBDB
Harpalus smaragdinus species	accepted (GBIF)	No associated record in PBDB
Harpalus smyrnensis species	accepted (GBIF)	No associated record in PBDB
Harpalus solitaris species	accepted (GBIF)	No associated record in PBDB
Harpalus somereni species	accepted (GBIF)	No associated record in PBDB
Harpalus somnulentus species	accepted (GBIF)	No associated record in PBDB
Harpalus spadiceus species	accepted (GBIF)	No associated record in PBDB
Harpalus spretus species	accepted (GBIF)	No associated record in PBDB
Harpalus spurius species	accepted (GBIF)	No associated record in PBDB
Harpalus stephani species	accepted (GBIF)	No associated record in PBDB
Harpalus stevenii species	accepted (GBIF)	No associated record in PBDB
Harpalus stierlini ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus stoetznerianus species	accepted (GBIF)	No associated record in PBDB
Harpalus strenuus species	accepted (GBIF)	No associated record in PBDB
Harpalus stricticollis species	accepted (GBIF)	No associated record in PBDB
Harpalus stygius ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus subaeneus species	accepted (GBIF)	No associated record in PBDB
Harpalus subcylindricus species	accepted (GBIF)	No associated record in PBDB
Harpalus subtruncatus species	accepted (GBIF)	No associated record in PBDB
Harpalus suensoni species	accepted (GBIF)	No associated record in PBDB
Harpalus sulphuripes species	accepted (GBIF)	No associated record in PBDB
Harpalus sundaicus species	accepted (GBIF)	No associated record in PBDB
Harpalus sushenicus species	accepted (GBIF)	No associated record in PBDB
Harpalus suturangulus species	accepted (GBIF)	No associated record in PBDB
Harpalus szalliesi species	accepted (GBIF)	No associated record in PBDB
Harpalus tabidus ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus taciturnus species	accepted (GBIF)	No associated record in PBDB
Harpalus tadorcus species	accepted (GBIF)	No associated record in PBDB
Harpalus tangutorum species	accepted (GBIF)	No associated record in PBDB
Harpalus tardigradus ✝ species	accepted (GBIF) listed (PBDB)		1	12.7 Ma Miocene	12.7 Ma Miocene
Harpalus tardus species	accepted (GBIF)	No associated record in PBDB
Harpalus tarsalis species	accepted (GBIF)	No associated record in PBDB
Harpalus tenebrosus species	accepted (GBIF)	No associated record in PBDB
Harpalus tenuestriatus species	accepted (GBIF)	No associated record in PBDB
Harpalus terrestris species	accepted (GBIF)	No associated record in PBDB
Harpalus texanus species	accepted (GBIF)	No associated record in PBDB
Harpalus tibeticus species	accepted (GBIF)	No associated record in PBDB
Harpalus tichonis species	accepted (GBIF)	No associated record in PBDB
Harpalus tinctulus species	accepted (GBIF)	No associated record in PBDB
Harpalus tiridates species	accepted (GBIF)	No associated record in PBDB
Harpalus tithonus species	accepted (GBIF)	No associated record in PBDB
Harpalus tjanschanicus species	accepted (GBIF)	No associated record in PBDB
Harpalus torosensis species	accepted (GBIF)	No associated record in PBDB
Harpalus torridoides species	accepted (GBIF)	No associated record in PBDB
Harpalus trichophorus species	accepted (GBIF)	No associated record in PBDB
Harpalus tridens species	accepted (GBIF)	No associated record in PBDB
Harpalus triseriatus species	accepted (GBIF)	No associated record in PBDB
Harpalus turcicus species	accepted (GBIF)	No associated record in PBDB
Harpalus turmalinus species	accepted (GBIF)	No associated record in PBDB
Harpalus udege species	accepted (GBIF)	No associated record in PBDB
Harpalus ulamaeformis species	accepted (GBIF)	No associated record in PBDB
Harpalus ulomaeformis ✝ species	accepted (GBIF) listed (PBDB)		1	37.2 Ma Eocene	37.2 Ma Eocene
Harpalus uniformis species	accepted (GBIF)	No associated record in PBDB
Harpalus ussuricus species	accepted (GBIF)	No associated record in PBDB
Harpalus ussuriensis species	accepted (GBIF)	No associated record in PBDB
Harpalus vagans species	accepted (GBIF)	No associated record in PBDB
Harpalus vanemdeni species	accepted (GBIF)	No associated record in PBDB
Harpalus venator species	accepted (GBIF)	No associated record in PBDB
Harpalus ventralis species	accepted (GBIF)	No associated record in PBDB
Harpalus vereschaginae species	accepted (GBIF)	No associated record in PBDB
Harpalus vernicosus species	accepted (GBIF)	No associated record in PBDB
Harpalus veterum ✝ species	accepted (GBIF) listed (PBDB)		1	50.3 Ma Eocene	50.3 Ma Eocene
Harpalus viridanus species	accepted (GBIF)	No associated record in PBDB
Harpalus viridicupreus species	accepted (GBIF)	No associated record in PBDB
Harpalus vittatus species	accepted (GBIF) listed (PBDB)		16	2.59 Ma Pleistocene	0 Ma Extant
Harpalus wadiensis species	accepted (GBIF)	No associated record in PBDB
Harpalus whitfieldi species	accepted (GBIF)	No associated record in PBDB
Harpalus whitfieldii ✝ species	listed (PBDB)		2	37.2 Ma Eocene	37.2 Ma Eocene
Harpalus wohlberedti species	accepted (GBIF)	No associated record in PBDB
Harpalus xanthopus species	accepted (GBIF)	No associated record in PBDB
Harpalus xinjiangensis species	accepted (GBIF)	No associated record in PBDB
Harpalus yinchuanensis species	accepted (GBIF)	No associated record in PBDB
Harpalus zabroides species	accepted (GBIF)	No associated record in PBDB
Harpalus zhdankoi species	accepted (GBIF)	No associated record in PBDB

Obsolete Names

Name	Source	Taxon Rank	Taxonomy
Brachypangus Tschitschérine, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Brachypangus*
Cephalomorphus Tschitscherin, 1897	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cephalomorphus*
Incisophonus Sciaky, 1986	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Incisophonus*
Plectralidus Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Plectralidus*
Micrarpalus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Micrarpalus*
Cordoharpalus Hatch, 1949	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cordoharpalus*
Macrophonus Tschitscherin, 1901	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Macrophonus*
Migadophonus Tschitscherin, 1897	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Migadophonus*
Lasioharpalus Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lasioharpalus*
Erpeinus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Erpeinus*
Opadius Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Opadius*
Pharalus Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pharalus*
Acardystus Reitter, 1908	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Acardystus*
Conicus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Conicus*
Pseudophonus Motschoulsky, 1848	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudophonus*
Harpalellus Lindroth, 1968	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpalellus*
Neoharpalus Mateu, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Neoharpalus*
Pangus Motschulsky, 1859	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pangus*
Platus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Platus*
Actephilus Stephens, 1833	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Actephilus*
Brachyophonus Sciaky, 1986	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Brachyophonus*
Hypsinephus Bates, 1878	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hypsinephus*
Pardileus Des Gozis, 1882	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pardileus*
Bioderus Motschoulsky, 1848	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Bioderus*
Typsarpalus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Typsarpalus*
Asmerinx Reitter, 1899	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Asmerinx*
Neopangus Tschitscherin, 1898	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Neopangus*
Pheuginus Motschoulsky, 1845	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pheuginus*
Parhalus Jeannel, 1946	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Parhalus*
Pseudoaphonus Chaudoir, 1863	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Pseudoaphonus*
Epharpalus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Epharpalus*
Nephoharpalus Huang, Lei, Yan & Hu, 1996	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Nephoharpalus*
Baeticoharpalus Serrano & Lencina, 2009	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Baeticoharpalus*
Crytophonus Brandmayr & Zetto-Brandmayr, 1982	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Crytophonus*
Acthephilus Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Acthephilus*
Megapangus Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Megapangus*
Metophonus Bedel, 1897	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Metophonus*
Neopardileus Habu, 1954	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Neopardileus*
Harpalobius Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpalobius*
Epiharpalus Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Epiharpalus*
Microderus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Microderus*
Empeirus Motschoulsky, 1844	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Empeirus*
Anisochirus Jeannel, 1946	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Anisochirus*
Loboharpalus Schauberger, 1932	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Loboharpalus*
Harpaloderus Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpaloderus*
Hesperophonus Antoine, 1959	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Hesperophonus*
Sulcophonus Schauberger, 1933	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Sulcophonus*
Harpaloxys Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpaloxys*
Lasiarpalus Jacobson, 1907	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Lasiarpalus*
Harpalomerus Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpalomerus*
Euharpalops Casey, 1924	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Euharpalops*
Amblystus Motschoulsky, 1864	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Amblystus*
Cephalotypsis Tschitscherin, 1901	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Cephalotypsis*
Microharpalus Tschitscherin, 1901	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Microharpalus*
Loxophonus Reitter, 1894	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Loxophonus*
Rapahlus Lucnik, 1922	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Rapahlus*
Ooistus Motschoulsky, 1864	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Ooistus*
Actophilus Agassiz, 1846	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Actophilus*
Glanodes Casey, 1914	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Glanodes*
Haploharpalus Schauberger, 1926	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Haploharpalus*
Harpaloxis Reitter, 1900	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Harpaloxis*
Artabas Gozis, 1882	GBIF	genus	Animalia : Arthropoda : Insecta : Coleoptera : Carabidae : *Artabas*

Synonymy List

Year	Name and Author
1802	Harpalus Latreille p. 92
1821	Harpalus Dejean p. 14
1861	Harpalus LeConte p. 33
1900	Harpalus Scudder p. 34
1917	Harpalus Wickham p. 464
1921	Harpalus Cockerell
2011	Harpalus Bouchard et al. p. 129
2011	Harpalus Tamutis et al.

References

Latreille P. A. , Histoire Naturelle, Générale et Particulière des Crustacés et des Insectes, Tome Troisième, 1-467

Dejean P. F. M. , Catalogue de la collection de Coléoptères de M. le Baron Dejean, 1-136

Scudder S. H. Adephagous and clavicorn Coleoptera from the Tertiary deposits at Florissant, Colorado with descriptions of a few other forms and a systematic list of the non-rhynchophorus Tertiary Coleoptera of North America, Monographs of the United States Geological Survey 40, 1-148

Wickham H. F. (1917) New Species of Fossil Beetles from Florissant, Colorado, Proceedings of the United States National Museum 52, 463-472

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Gustafsson, B. (Col.) (2004) Catalogus Coleopterorum Sueciae. Naturhistoriska riksmuseet, Stockholm, Excelfil hämtad från http://www.nrm.se/en/catalogus.html.se. - via Dyntaxa. Svensk taxonomisk databas

CoL2006 - via The Interim Register of Marine and Nonmarine Genera

Hallan, J. (2000-current). Biology Catalog. Web compilation accessible at http://bug.tamu.edu/research/collection/hallan/ (accessed June 2012). - via The Interim Register of Marine and Nonmarine Genera

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Nomenclator Zoologicus. A list of the names of genera and subgenera in zoology from the tenth edition of Linnaeus, 1758 to the end of 2004. Digitised by uBio from vols. 1-9 of Neave (ed.), 1939-1996 plus supplementary digital-only volume. http://ubio.org/NomenclatorZoologicus (as at 2006). - via The Interim Register of Marine and Nonmarine Genera

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as per family (nonmarine except where stated) - via The Interim Register of Marine and Nonmarine Genera

Noonan, Gerald R., 1991: null. Classification, Cladistics, and Natural History of Native North American Harpalus Latreille (Insecta: Coleoptera: Carabidae: Harpalini), Excluding Subgenera Glanodes and Pseudophonus. viii + 310. - via Integrated Taxonomic Information System (ITIS)

Bousquet, Yves, and André Larochelle, 1993: Catalogue of the Geadephaga (Coleoptera: Trachypachidae, Rhysodidae, Carabidae including Cicindelini) of America North of Mexico. Memoirs of the Entomological Society of Canada, no. 167. 397. - via Integrated Taxonomic Information System (ITIS)

Bousquet, Yves, 2012: Catalogue of Geadephaga (Coleoptera, Adephaga) of America, north of Mexico. ZooKeys, issue 245. 1-1722. - via Integrated Taxonomic Information System (ITIS)

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

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