Quick Links : Mindat手册 The Rock H. Currier Digital Library Mindat Newsletter [Free Download]

主页关于 Mindat Mindat手册 Mindat的历史版权 Who We Are 联系我们于 Mindat.org刊登广告

捐赠给 Mindat Corporate Sponsorship 赞助板页已赞助的板页在 Mindat刊登广告的广告商于 Mindat.org刊登广告

Learning Center What is a mineral?The most common minerals on earth Information for Educators Mindat Articles The Elements The Rock H. Currier Digital Library Geologic Time

搜索矿物的性质搜索矿物的化学 Advanced Locality Search 随意显示任何一种矿物 Random Locality 使用minID搜索邻近产地 Search Articles 搜索词汇表更多搜索选项

Mindat手册添加新照片 Rate Photos 产区编辑报告 Coordinate Completion Report 添加词汇表项目

Mining Companies 统计会员列表 Mineral Museums Clubs & Organizations 矿物展及活动 The Mindat目录表设备设置 The Mineral Quiz

照片搜索 Photo Galleries Search by Color 今天最新的照片昨天最新的照片用户照片相集过去每日精选照片相集 Photography

BETA TEST - Fossil data and pages are very much experimental and under development. Please report any problems

Varanoidea

Photos

Photo Gallery

Source Data

Source	ID	Link
Global Biodiversity Information Facility ID (GBIF)	Not linked to a GBIF record.
PaleoBioDB ID (PBDB)	38023	https://paleobiodb.org/classic/checkTaxonInfo?taxon_no=38023

Rank

superfamily (PBDB)

Taxonomy (PBDB)

Life : Animalia : Chordata : Reptilia : Squamata : Varanoidea

Classification
(PBDB,GBIF)

Rank	Name	Author
-	Eukaryota
-	Opisthokonta	Cavalier-Smith 1987
kingdom	Animalia
-	Triploblastica	Lankester 1877
-	Nephrozoa	Jondelius et al. 2002
-	Deuterostomia	Grobben 1908
phylum	Chordata	Haeckel 1847
subphylum	Vertebrata	Lamarck 1801
superclass	Gnathostomata	Gegenbauer 1874
-	Osteichthyes
-	Sarcopterygii	Romer 1955
subclass	Dipnotetrapodomorpha
subclass	Tetrapodomorpha
-	Tetrapoda
-	Reptiliomorpha	Säve-Söderbergh 1934
-	Anthracosauria	Säve-Söderbergh 1934
-	Batrachosauria
-	Cotylosauria
-	Amniota	Haeckel 1866
-	Sauropsida	Huxley 1864
class	Reptilia	Laurenti 1768
subclass	Eureptilia
-	Romeriida	Gauthier et al. 1988
-	Diapsida
-	Eosuchia
-	Neodiapsida	Benton 1985
-	Sauria	Gauthier 1984
-	Lepidosauromorpha
superorder	Lepidosauria
order	Squamata	Oppel 1811
-	Episquamata	Pyron et al. 2013
-	Toxicofera	Vidal and Hedges 2005
suborder	Anguimorpha	Fürbringer 1900
-	Anguiformes	Conrad 2006
infraorder	Platynota	Baur 1890
superfamily	Varanoidea	Gray 1827

Opinions (PBDB)

Name	Rank	Opinion	Evidence	Author
Varanoidea	superfamily	belongs to Platynota	stated without evidence	Baur, 1890
Varanoidea	superfamily	belongs to Anguimorpha	stated with evidence	Camp, 1923
Varanoidea	superfamily	subjective synonym of Platynota	stated with evidence	McDowell and Bogert, 1954
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Antunes, 1964
Varanoidea	superfamily	belongs to Platynota	stated with evidence	Estes, 1964
Varanoidea	superfamily	belongs to Anguimorpha	stated with evidence	Kuhn, 1966
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Seiffert, 1975
Varanoidea	superfamily	belongs to Anguimorpha	stated with evidence	Estes, 1983
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Stritzke, 1983
Varanoidea	superfamily	belongs to Diploglossa	implied	Carroll, 1988
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Gao and Fox, 1996
Varanoidea	superfamily	belongs to Squamata	stated without evidence	Lee and Caldwell, 2000
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Peng et al., 2001
Varanoidea	superfamily	belongs to Platynota	stated with evidence	Conrad, 2006
Varanoidea	superfamily	belongs to Platynota	stated without evidence	Norell et al., 2007
Varanoidea	superfamily	belongs to Anguimorpha	stated without evidence	Houssaye et al., 2013
Varanoidea	superfamily	belongs to Reptilia	stated without evidence	Houssaye et al., 2013

Status (PBDB)

extant

Taxon Size (PBDB)

Extant Size (PBDB)

35 (49%)

First Recorded Appearance

130 - 122 Ma
Early/Lower Cretaceous

Environment

terrestrial (based on Diapsida)

Motility

actively mobile (based on Osteichthyes)

Diet

carnivore (based on Varanoidea)

Reproduction

oviparous (based on Varanoidea)

Taphonomy

phosphatic (based on Vertebrata)

Primary Reference (PBDB)

R. L. Carroll. 1988. Vertebrate Paleontology and Evolution

Subtaxa

Name	Status	Common Name(s)	Fossil Occurrences	Oldest	Youngest
Helodermatidae family	accepted (GBIF) listed (PBDB)		26	83.6 Ma Late/Upper Cretaceous	0 Ma Extant
Lanthanotidae family	accepted (GBIF) listed (PBDB)				0 Ma Extant
Pachyvaranidae ✝ family	listed (PBDB)		9	83.5 Ma Late/Upper Cretaceous	70.6 Ma Late/Upper Cretaceous
Parasaniwidae ✝ family	accepted (GBIF) listed (PBDB)		2	93.5 Ma Late/Upper Cretaceous	58.7 Ma Paleocene
Varanidae family	accepted (GBIF) listed (PBDB)	Monitor Lizard	118	86.3 Ma Late/Upper Cretaceous	0 Ma Extant
Saniwinae ✝ subfamily	listed (PBDB)		69	63.3 Ma Paleocene	33.9 Ma Oligocene
Aiolosaurus ✝ genus	accepted (GBIF) listed (PBDB)		1	83.6 Ma Late/Upper Cretaceous	83.6 Ma Late/Upper Cretaceous
Arcanosaurus ✝ genus	accepted (GBIF) listed (PBDB)		1	130 Ma Early/Lower Cretaceous	130 Ma Early/Lower Cretaceous
Heloderma genus	accepted (GBIF) listed (PBDB)	Beaded Lizard	5	24.8 Ma Oligocene	0 Ma Extant
Judeasaurus ✝ genus	accepted (GBIF) listed (PBDB)		1	99.6 Ma Late/Upper Cretaceous	99.6 Ma Late/Upper Cretaceous
Ovoo ✝ genus	accepted (GBIF) listed (PBDB)		1	83.6 Ma Late/Upper Cretaceous	83.6 Ma Late/Upper Cretaceous

Synonymy List

Year	Name and Author
1827	Varanoidea Gray
1890	Helodermatoidea Baur
1890	Varanoidea Baur p. 262
1923	Varanoidea Camp
1964	Varanoidea Antunes p. 137
1964	Varanoidea Estes p. 128
1966	Varanoidea Kuhn p. 56
1975	Varanoidea Seiffert p. 10
1983	Varanoidea Estes

Year	Name and Author
1983	Varanoidea Stritzke
1988	Varanoidea Carroll
1996	Varanoidea Gao and Fox p. 88
2000	Varanoidea Lee and Caldwell
2001	Varanoidea Peng et al. p. 32
2006	Varanoidea Conrad p. 122
2007	Varanoidea Norell et al. p. 3
2013	Varanoidea Houssaye et al.
2013	Varanoidea Houssaye et al.

References

Carroll R. L. (1988) , Vertebrate Paleontology and Evolution, 1-698

Antunes M. T. (1964) O Neocretacio e o Cenozoico do litoral de Angola, Junta de Investigcoes do Ultramar, 1-254

Estes R. (1964) Fossil vertebrates from the Late Cretaceous Lance Formation, eastern Wyoming, University of California Publications in Geological Sciences 49, 1-187

Kuhn O. W. M. (1966) , Die Reptilien. System und Stammesgeschichte [The Reptiles. Systematics and Phylogeny.], 1-154

Seiffert J. (1975) Upper Jurassic lizards from central Portugal, Contribuição para o conhecimento da Fauna do Kimerridgiano da Mina de Lignito Guimarota (Leiria, Portugal). Serviços Geológicos de Portugal, Memória (Nova Série) 22, 7-85

Estes R. (1983) Sauria terrestria, Amphisbaenia, Handbuch der Paläoherpetologie 10A, 1-249

Stritzke R. (1983) Saniwa feisti n. sp., ein Varanide (Lacertilia, Reptilia) aus dem Mittel-Eozän von Messel bei Darmstadt, Senckenbergiana Lethaea 64 5/6, 497-508

Gao K. -Q., Fox R. C. (1996) Taxonomy and evolution of Late Cretaceous lizards (Reptilia: Squamata) from western Canada, Bulletin of the Carnegie Museum of natural History 33, 1-107

Lee M. S. Y., Caldwell M. W. (2000) Adriosaurus and the affinities of mosasaurs, dolichosaurs, and snakes, Journal of Paleontology 74 3, 915-937

Peng J., Russell A. P., et al (2001) Vertebrate microsite assemblages (exclusive of mammals) from the Foremost and Oldman Formations of the Judith River Group (Campanian) of southeastern Alberta: an illustrated guide, Provincial Museum of Alberta, Natural History Occasional Paper 25, 1-54

Conrad J. L. (2006) An Eocene shinisaurid (Reptilia, Squamata) from Wyoming, U.S.A., Journal of Vertebrate Paleontology 26 1, 113-126

Norell M. A., Gao K.-Q., et al (2007) A New Platynotan Lizard (Diapsida: Squamata) from the Late Cretaceous Gobi Desert (Ömnögov), Mongolia, American Museum Novitates 3605, 1-22

Houssaye A., Bardet N., et al (2013) Squamate finding in "Lo Hueco" (Late Campanian-Early Maastrichtian, Cuenca Province, Spain): the second non-marine pythonomorph lizard, Palaeontologische Zeitschrift 87 3, 415-422 doi:10.1007/s12542-013-0164-6

Houssaye A., Rage J.-C., et al (2013) A new varanoid squamate from the Early Cretaceous (Barremian-Aptian) of Burgos, Spain, Cretaceous Research 41, 127-135 doi:10.1016/j.cretres.2012.11.005

Data courtesy of: PBDB: The Paleobiology Database, Creative Commons CC-BY licenced. , GBIF: the Global Biodiversity Information Facility, various licences, iDigBio, various licences, and EOL: The Encyclopedia of Life (Open Data Public Domain). Because fossils are made of minerals too!

Quick NavTop Subtaxa Synonymy List References

Go to top of page